Luolishaniidae
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Luolishaniidae
The Luolishaniidae or Luolishaniida are a group of Cambrian and Ordovician lobopodians with anterior 5 or 6 pairs of setiferous lobopods. Most luolishaniids also have posterior lobopods each with a hooked claws, and thorn-shaped sclerites arranged as three or more per trunk segment. The type genus is based on ''Luolishania longicruris'' Hou and Chen, 1989, from the Chengjiang Lagerstatte, South China. They are presumed to have been benthic suspension or filter feeders. New specimens of the previously enigmatic ''Facivermis ''Facivermis'' (meaning "torch worm" ) is a genus of sessile lobopodian from the Lower Cambrian Maotianshan shales of China Anatomy ''Facivermis'' was a worm-like creature up to 90 mm long. Its body was divided into three sections. The ante ...'' show that it was a sessile tube-dweller, and part of this group. References Lobopodia Cambrian invertebrates Prehistoric protostome families {{paleo-protostome-stub ...
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Collinsovermis
''Collinsovermis'' is a genus of extinct panarthropod belonging to the group Lobopodia and known from the middle Cambrian Burgess Shale in British Columbia, Canada. It is monotypic having only one species, ''Collinsovermis monstruosus''. After its initial discovery in 1983, Desmond H. Collins popularised it as a unique animal and was subsequently dubbed "Collins' monster" for its unusual super armoured body. The formal scientific description and name were given in 2020. Discovery ''Collinsovermis'' was discovered in 1983 by Desmond H. Collins, curator of invertebrate palaeontology at the Royal Ontario Museum, from an expedition at Mount Stephen at the Yoho National Park, British Columbia, Canada. It was found among the Burgess Shale that belonged to the middle Cambrian period called Wuliuan, which is around 509 to 505 million years ago. In 1985, Collins presented the discovery before the Geological Society of America, and published the next year in a popular magazine ''Rotunda'' ...
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Luolishania
''Luolishania'' is an extinct genus of lobopodian worm and known from the Lower Cambrian Chiungchussu Formation (Maotianshan Shales) of the Chengjiang County, Yunnan Province, China. A monotypic genus, it contains one species ''Luolishania longicruris''. It was discovered and described by Hou Xian-Guang and Chen Jun-Yuan in 1989. It is one of the superarmoured Cambrian lobopodians suspected to be either an intermediate form in the origin of velvet worms (Onychophora) or basal to at least Tardigrada and Arthropoda. It is the basis of the family name Luolishaniidae, which also include other related lobopods such as '' Acinocricus'', ''Collinsium'', ''Facivermis,'' and ''Ovatiovermis''. Along with ''Microdictyon'', it is the first lobopodian fossil discovered from China. Discovery A single specimen of ''Luolishania'' fossil was discovered by Hou Xian-guang and Chen Jun-yuan of the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, from the Chengjiang Lage ...
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Ovatiovermis
''Ovatiovermis'' is a genus of filter-feeding lobopodian known from the Burgess Shale. Like many lobopodians, it had nine pairs of lobopods (legs). It was well adapted to filter-feeding and probably did so from the nearest high vantage point. Morphology ''Ovatiovermis'' had nine pairs of lobopods. The first two pairs were elongate and had approximately 20 pairs of spines on each lobopod, with a bifid claw at each tip. The third to sixth pairs of lobopods were shorter and their paired spines were much smaller. On these four pairs of lobopods the spines were only large near the tip. The last three pairs of lobopods did not have the paired spines, showing that these spines were used for filter-feeding. Instead these lobopods had hooked claws with which it could grip one of the corals or sponges found at the time and rear up into the current. At the anterior end of the animal it had an evertible proboscis, with its mouth at the end. This would probably have been used to suck any ...
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Facivermis
''Facivermis'' (meaning "torch worm" ) is a genus of sessile lobopodian from the Lower Cambrian Maotianshan shales of China Anatomy ''Facivermis'' was a worm-like creature up to 90 mm long. Its body was divided into three sections. The anterior section had five equally sized pairs of appendages with two setal rows along the margins. The middle section was elongate and five times longer than the anterior or posterior. The posterior section was pear-shaped and had three rows of hooks surrounding the anus. Classification ''Facivermis'' was considered by its describers to be a polychaete worm. An affinity to the unusual crustacean lineage Pentastomida has also been proposed, but is seen as unlikely. Since its discovery, however, most evidence has supported its being a lobopodian. Liu ''et al.'' draw a comparison to the known lobopodian '' Miraluolishania''. Liu ''et al.'' also note that the pear-shaped end bears a close resemblance to the proboscis of priapulid worms if it is i ...
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Acinocricus
''Acinocricus'' is a genus of extinct worm belonging to the group Lobopodia and known from the middle Cambrian Spence Shale of Utah, United States. As a monotypic genus, it has one species ''Acinocricus stichus''. The only lobopodian discovered from the Spence Shale, it was described by Simon Conway Morris and Richard A. Robison in 1988. Owing to the original fragmentary fossils discovered since 1982, it was initially classified as an alga, but later realised to be an animal belonging to Cambrian fauna. Discovery The first specimen of ''Acinocricus'' was discovered by American palaeontologist Lloyd Gunther in 1982 from the Spence Shale in Miners Hollow, Wellsville Mountains, Utah. It was embedded in hardened mud and was incomplete with some of its body part missing. More than a dozen fragmentary fossils were later recovered from the same site and the surrounding areas. Simon Conway Morris of the University of Cambridge and Richard A. Robison of the University of Kansas jointly p ...
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Miraluolishania
''Miraluolishania'' is an extinct lobopodian known from Chengjiang County in China. It is remarkable for the possession of lensed pit-eyes. The only species, ''Miraluolishania haikouensis'', was described from the Maotianshan Shales at Haikou by Jianni Liu and Degan Shu in 2004. In 2009, a team of palaeontologists at the Yunnan University, led by Xiaoya Ma reported the discovery of 42 other specimens from Haikou. With the help of Swiss palaeontologist Jan Bergström, Ma and Hou came to the conclusion that all the specimens were the same species (junior synonym) as ''Luolishania ''Luolishania'' is an extinct genus of lobopodian worm and known from the Lower Cambrian Chiungchussu Formation (Maotianshan Shales) of the Chengjiang County, Yunnan Province, China. A monotypic genus, it contains one species ''Luolishania longic ...'';'''' another lobopod discovered from the Chengjian in 1989. Chengjian is 40 kms from Haikou and the fossil fauna are different. A reassessment by Liu and S ...
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Collinsium
''Collinsium'' was a genus of lobopodian from the Early Cambrian. It is represented by a single fossil species, ''Collinsium ciliosum'', found in the Xiaoshiba Lagerstätte ( Hongjingshao Formation) of China. Similar to the later ''Hallucigenia'' it was a small worm-like creature with spikes along its back and feeding tentacles near its head. Unlike ''Hallucigenia'', ''Collinsium'' had 9 walking appendages and 4 fine anterior appendages. Its body was covered in hair-like Papillae, and its fine anterior appendages were lined with Setae likely used for filter feeding. The generic name honors paleontologist Desmond Collins Desmond H. Collins is a Canadian paleontologist, associate professor of zoology at the University of Toronto The University of Toronto (UToronto or U of T) is a public university, public research university in Toronto, Ontario, Canada, locat .... References {{Taxonbar, from=Q20647066 Prehistoric protostome genera Cambrian China Fossils of China Fo ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Lobopodian
The lobopodians, members of the informal group Lobopodia (from the Greek language, Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as ''Aysheaia'' and ''Hallucigenia''. The oldest near-complete fossil lobopodians date to the Lower Cambrian; some are also known from Ordovician, Silurian and Carboniferous Lagerstätten. Some bear toughened claws, plates or spines, which are commonly preserved as small carbonaceous fossil, carbonaceous or Small Shelly Fossils, mineralized microfossils in Cambrian strata. The grouping is considered to be paraphyletic, as the three living panarthropod groups (Arthropoda, Tardigrada and Onychophora) are thought to have evolved from lobopodian ...
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Maotianshan Shales
The Maotianshan Shales are a series of Cambrian, Early Cambrian deposits in the Chiungchussu Formation, famous for their ''Lagerstätte, Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale. They take their name from Maotianshan Hill (, Literal meaning: Hat Sky Mountain) in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. The shales also contain the slightly younger Guanshan biota from Mal ...
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