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Lar1
LAR1 ('Lichen-Associated Rhizobiales 1') refers to a specific bacterial lineage in the order Hyphomicrobiales (formerly Rhizobiales) that has most frequently been found directly in association with lichens. This lineage is currently known to associate with lichens that have a green-algal photosynthetic partner (as opposed to a cyanobacterial partner) and a fungal partner in the Lecanoromycetes (though other groups of fungi have not yet been examined). This lineage has been documented in association with all green-algal lichens specifically tested (all from North America), and was also found in a sequence library derived from Antarctic lichens. The specific ecological niche occupied by this lineage indicates that it may rely on certain nutrients that are abundant in green-algal lichen thalli but are rarer in other environments. Nitrogen fixing The LAR1 lineage is currently defined based on sequences of the 16S rRNA gene 16S or 16s may refer to: * Ribosomal RNAs, in biology: * ...
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Hyphomicrobiales
The ''Hyphomicrobiales'' are an order of Gram-negative Alphaproteobacteria. The rhizobia, which fix nitrogen and are symbiotic with plant roots, appear in several different families. The four families ''Nitrobacteraceae'', ''Hyphomicrobiaceae'', '' Phyllobacteriaceae'', and ''Rhizobiaceae'' contain at least several genera of nitrogen-fixing, legume-nodulating, microsymbiotic bacteria. Examples are the genera ''Bradyrhizobium'' and ''Rhizobium''. Species of the ''Methylocystaceae'' are methanotrophs; they use methanol (CH3OH) or methane (CH4) as their sole energy and carbon sources. Other important genera are the human pathogens ''Bartonella'' and ''Brucella'', as well as ''Agrobacterium'' (useful in genetic engineering). Taxonomy Accepted families * ''Aestuariivirgaceae'' Li ''et al''. 2019 * '' Afifellaceae'' Hördt ''et al''. 2020 * '' Ahrensiaceae'' Hördt ''et al''. 2020 * ''Alsobacteraceae'' Sun ''et al''. 2018 * ''Amorphaceae'' Hördt ''et al''. 2020 * ''Ancalomicrobia ...
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16S Ribosomal RNA
16 S ribosomal RNA (or 16 S rRNA) is the RNA component of the 30S subunit of a prokaryotic ribosome (SSU rRNA). It binds to the Shine-Dalgarno sequence and provides most of the SSU structure. The genes coding for it are referred to as 16S rRNA gene and are used in reconstructing phylogenies, due to the slow rates of evolution of this region of the gene. Carl Woese and George E. Fox were two of the people who pioneered the use of 16S rRNA in phylogenetics in 1977. Multiple sequences of the 16S rRNA gene can exist within a single bacterium. Functions * Like the large (23S) ribosomal RNA, it has a structural role, acting as a scaffold defining the positions of the ribosomal proteins. * The 3-end contains the anti- Shine-Dalgarno sequence, which binds upstream to the AUG start codon on the mRNA. The 3-end of 16S RNA binds to the proteins S1 and S21 which are known to be involved in initiation of protein synthesis * Interacts with 23S, aiding in the binding of the two ribosomal s ...
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Horizontal Gene Transfer
Horizontal gene transfer (HGT) or lateral gene transfer (LGT) is the movement of genetic material between Unicellular organism, unicellular and/or multicellular organisms other than by the ("vertical") transmission of DNA from parent to offspring (reproduction). HGT is an important factor in the evolution of many organisms. HGT is influencing scientific understanding of higher order evolution while more significantly shifting perspectives on bacterial evolution. Horizontal gene transfer is the primary mechanism for the spread of antibiotic resistance in bacteria, and plays an important role in the evolution of bacteria that can degrade novel compounds such as human-created Bactericide, pesticides and in the evolution, maintenance, and transmission of virulence. It often involves Temperateness (virology), temperate bacteriophages and plasmids. Genes responsible for antibiotic resistance in one species of bacteria can be transferred to another species of bacteria through various m ...
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The ISME Journal
''The ISME Journal: Multidisciplinary Journal of Microbial Ecology'' is a peer-reviewed scientific journal that covers diverse and integrated areas of microbial ecology spanning the breadth of microbial life, including bacteria, archaea, microbial eukaryotes, and viruses. It is an official publication of the International Society for Microbial Ecology (ISME) and publishes original research articles, reviews, and commentaries. The founding editors-in-chief are Mark Bailey and George Kowalchuk. The journal is published on behalf of ISME by the Nature Publishing Group. According to the ''Journal Citation Reports'' published in 2021, the journal had an impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate that reflects the yearly mean number of citations of articles published in the last two years in a given journal, as ... of 10.302. The SCImago Journal Rank placed the ISME Journal 9th ou ...
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Nif Gene
The ''nif'' genes are genes encoding enzymes involved in the fixation of atmospheric nitrogen into a form of nitrogen available to living organisms. The primary enzyme encoded by the ''nif'' genes is the nitrogenase complex which is in charge of converting atmospheric nitrogen (N2) to other nitrogen forms such as ammonia which the organism can use for various purposes. Besides the nitrogenase enzyme, the ''nif'' genes also encode a number of regulatory proteins involved in nitrogen fixation. The ''nif'' genes are found in both free-living nitrogen-fixing bacteria and in symbiotic bacteria associated with various plants. The expression of the ''nif'' genes is induced as a response to low concentrations of fixed nitrogen and oxygen concentrations (the low oxygen concentrations are actively maintained in the root environment of host plants). The first Rhizobium genes for nitrogen fixation (nif) and for nodulation (nod) were cloned in the early 1980s by Gary Ruvkun and Sharon R. Long i ...
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Activation Energy
In chemistry and physics, activation energy is the minimum amount of energy that must be provided for compounds to result in a chemical reaction. The activation energy (''E''a) of a reaction is measured in joules per mole (J/mol), kilojoules per mole (kJ/mol) or kilocalories per mole (kcal/mol). Activation energy can be thought of as the magnitude of the potential barrier (sometimes called the energy barrier) separating minima of the potential energy surface pertaining to the initial and final thermodynamic state. For a chemical reaction to proceed at a reasonable rate, the temperature of the system should be high enough such that there exists an appreciable number of molecules with translational energy equal to or greater than the activation energy. The term "activation energy" was introduced in 1889 by the Swedish scientist Svante Arrhenius. Other uses Although less commonly used, activation energy also applies to nuclear reactions and various other physical phenomena. Te ...
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Nitrogen
Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at seventh in total abundance in the Milky Way and the Solar System. At standard temperature and pressure, two atoms of the element bond to form N2, a colorless and odorless diatomic gas. N2 forms about 78% of Earth's atmosphere, making it the most abundant uncombined element. Nitrogen occurs in all organisms, primarily in amino acids (and thus proteins), in the nucleic acids ( DNA and RNA) and in the energy transfer molecule adenosine triphosphate. The human body contains about 3% nitrogen by mass, the fourth most abundant element in the body after oxygen, carbon, and hydrogen. The nitrogen cycle describes the movement of the element from the air, into the biosphere and organic compounds, then back into the atmosphere. Many indus ...
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Nitrogen Fixation
Nitrogen fixation is a chemical process by which molecular nitrogen (), with a strong triple covalent bond, in the air is converted into ammonia () or related nitrogenous compounds, typically in soil or aquatic systems but also in industry. Atmospheric nitrogen is molecular dinitrogen, a relatively nonreactive molecule that is metabolically useless to all but a few microorganisms. Biological nitrogen fixation or ''diazotrophy'' is an important microbials mediated process that converts dinitrogen (N2) gas to ammonia (NH3) using the nitrogenase protein complex (Nif). Nitrogen fixation is essential to life because fixed inorganic nitrogen compounds are required for the biosynthesis of all nitrogen-containing organic compounds, such as amino acids and proteins, nucleoside triphosphates and nucleic acids. As part of the nitrogen cycle, it is essential for agriculture and the manufacture of fertilizer. It is also, indirectly, relevant to the manufacture of all nitrogen chemical c ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Ecological Niche
In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for example, by growing when resources are abundant, and when predators, parasites and pathogens are scarce) and how it in turn alters those same factors (for example, limiting access to resources by other organisms, acting as a food source for predators and a consumer of prey). "The type and number of variables comprising the dimensions of an environmental niche vary from one species to another ndthe relative importance of particular environmental variables for a species may vary according to the geographic and biotic contexts". See also Chapter 2: Concepts of niches, pp. 7 ''ff'' A Grinnellian niche is determined by the habitat in which a species lives and its accompanying behavioral adaptations. An Eltonian niche emphasizes that a spec ...
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Lecanoromycetes
Lecanoromycetes is the largest class of lichenized fungi. It belongs to the subphylum Pezizomycotina in the phylum Ascomycota. The asci (spore-bearing cells) of the Lecanoromycetes most often release spores by rostrate dehiscence. Genera of uncertain placement The are several genera in the Lecanoromycetes that have not been placed into any order or family. These are: *''Argopsis'' – 1 sp. *''Ascographa ''Ascographa'' is a genus of fungi in the Helotiales order. The relationship of this taxon to other taxa within the order is unknown (''incertae sedis''), and it has not yet been placed with certainty into any family. This is a monotypic In b ...'' - 1 sp. *'' Bartlettiella'' – 1 sp. *'' Bouvetiella'' – 1 sp. *'' Buelliastrum'' – 1 sp. *'' Haploloma'' – 1 sp. *'' Hosseusia'' – 3 spp. *'' Korfiomyces'' – 1 sp. *'' Maronella'' – 1 sp. *'' Notolecidea'' – 1 sp. *'' Petractis'' – 3 spp. *'' Piccolia'' – 10 spp. *'' Ravenelula'' – 2 spp. *'' R ...
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