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Loxommatinae
Baphetidae is an extinct family of early tetrapods. Baphetids were large labyrinthodont predators of the Late Carboniferous period ( Namurian through Westphalian) of Europe. Fragmentary remains from the Early Carboniferous of Canada have been tentatively assigned to the group. The phylogenetic relationships of baphetids is uncertain; while many studies have placed the group as a close relative of Amniota, other analyses have found Baphetidae to be a more basal clade of early stem tetrapods. Baphetids were among the first of the Carboniferous fossil tetrapods to be found and were originally described in 1850 by John William Dawson. The baphetids have been referred to the family Loxommatidae, but this group was later shown to be a junior synonym of Baphetidae, which was named earlier in 1865. Baphetids are known mainly from skulls; very little postcranial material has been found. Palaeobiology The presence of lateral lines and the long rows of needle-like teeth show that most wer ...
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Loxomma
''Loxomma'' (meaning “slanting eyes”) is an extinct genus of Loxommatinae and one of the first Carboniferous tetrapods. They were first described in 1862 and further described in 1870 when two more craniums were found. It is mostly associated with the area of the United Kingdom. They share features with modern reptiles as well as with fish. They had 4 paddle-like limbs that they used to swim in lakes, but they breathed air. Their diet consisted mostly of live fish. They are of the family Baphetidae which are distinguished by their keyhole shaped orbits, while ''Loxomma'' themselves are distinguished by the unique texture on their skulls, said to be honeycomb-like. History and discovery The genus ''Loxomma'' was discovered and named by Thomas Henry Huxley in 1862 via a near perfect cranium, a vertebra and a rib in the Lanarkshire coal-field of Scotland. The family Baphetidae were among the first Carboniferous tetrapods to be found and were first described by William Da ...
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Kyrinion
''Kyrinion'' is an extinct genus of baphetid tetrapod from the Late Carboniferous of England. It is known from a skull that was found in Tyne and Wear county dating back to the Westphalian (stage), Westphalian stage. Along with the skull is part of the lower jaw, an arch of the Atlas (anatomy), atlas bone (the vertebra that connects to the skull) and a rib possibly belonging to a cervical (neck) vertebra. The type species ''K. martilli'' was named from this material in 2003. Description The holotype skull of ''Kyrinion'' is well preserved with the back of the skull and both stapes, or ear bones, intact. The orbit (anatomy), orbits, or eye sockets, are somewhat triangular in shape. Areas connecting the palate with the jaws and braincase make the skull inflexible. Lateral lines, used for sensory perception in aquatic environments, are present on the quadratojugal bone behind the eyes. The lower jaw lacks the distinctive features of some other baphetids, such as teeth on the paras ...
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Spathicephalus
''Spathicephalus'' is an extinct genus of stem tetrapods (early four-limbed vertebrates) that lived during the middle of the Carboniferous Period. The genus includes two species: the type species ''S. mirus'' from Scotland, which is known from two mostly complete skulls and other cranial material, and the species ''S. pereger'' from Nova Scotia, which is known from a single fragment of the skull table. Based on the ''S. mirus'' material, the appearance of ''Spathicephalus'' is unlike that of any other early tetrapod, with a flattened, square-shaped skull and jaws lined with hundreds of very small chisel-like teeth. However, ''Spathicephalus'' shares several anatomical features with a family of stem tetrapods called Baphetidae, leading most paleontologists who have studied the genus to place it within a larger group called Baphetoidea, often as part of its own monotypic family Spathicephalidae. ''Spathicephalus'' is thought to have fed on aquatic invertebrates through a combinati ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Amniote
Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are distinguished from the other tetrapod clade — the amphibians — by the development of three extraembryonic membranes ( amnion for embryoic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage), thicker and more keratinized skin, and costal respiration (breathing by expanding/constricting the rib cage). All three main features listed above, namely the presence of an amniotic buffer, water-impermeable cutes and a robust respiratory system, are very important for amniotes to live on land as true terrestrial animals – the ability to reproduce in locations away from water bodies, better homeostasis in drier environments, and more efficient air respiration to power terrestrial locomotions, although the ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Batrachosauria
Batrachosauria ("Frog Saurians") is a name given either to very reptile-like amphibians dating from the Carboniferous and Permian periods, or to amniotes and those amphibians very closely related to them. In current cladistic schemes, Batrachosauria is the sister clade to the Anthracosauroidea Characteristics From the Palaeos website: * Intercentrum reduced * large canine-like maxillary tooth See also * Reptiliomorpha Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was def ... References External links * PalaeosBatrachosauria Reptiliomorphs {{paleo-amphibian-stub ...
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Richard Lydekker
Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, a barrister-at-law with Dutch ancestry. The family moved to Harpenden Lodge soon after Richard's birth. He was educated at Trinity College, Cambridge, where he took a first-class in the Natural Science tripos (1872). In 1874 he joined the Geological Survey of India and made studies of the vertebrate palaeontology of northern India (especially Kashmir). He remained in this post until the death of his father in 1881. His main work in India was on the Siwalik palaeofauna; it was published in ''Palaeontologia Indica''. He was responsible for the cataloguing of the fossil mammals, reptiles, and birds in the Natural History Museum (10 vols., 1891). He named a variety of taxa including the golden-bellied mangabey; as a taxon authority he is nam ...
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Crassigyrinus
''Crassigyrinus'' (from la, crassus, 'thick' and el, γυρίνος el, gyrínos, 'tadpole') is an extinct genus of carnivorous stem tetrapod from the Early Carboniferous Limestone Coal Group of Scotland and possibly Greer, West Virginia. The type specimen was originally described as ''Macromerium scoticum'' and lacked a complete skull. With subsequent discoveries, ''Crassigyrinus'' is now known from three skulls, one of which is in articulation with a fairly complete skeleton, and two incomplete lower jaws. ''Crassigyrinus'' grew up to 2 meters in length, coupled with tiny limbs and unusually large jaws. ''Crassigyrinus'' is taxonomically enigmatic, having confused paleontologists for decades with its apparent fish-like and tetrapod features.
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Eucritta
''Eucritta'' (meaning "true creature") is an extinct genus of stem-tetrapod from the Viséan epoch in the Carboniferous period of Scotland. The name of the type and only species, ''E. melanolimnetes'' ("true creature from the black lagoon") is a homage to the 1954 horror film ''Creature from the Black Lagoon''. ''Eucritta'' possessed many features in common with other generalized Carboniferous tetrapods and tetrapod relatives. A large amount of these features were plesiomorphic, meaning that they resembled the "primitive" condition that was acquired when four-limbed vertebrates ("amphibians", in the broad sense) first appeared. With a short, wide skull, large eyes, and strongly-built limbs, ''Eucritta'' proportionally resembled ''Balanerpeton'', a contemporary terrestrial tetrapod which was one of the earliest members of Temnospondyli, a successful tetrapod group which may have produced modern amphibians. However, ''Eucritta'' lacked key temnospondyl adaptations, nor certain adap ...
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Dentary
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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