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Laying Worker Bee
A laying worker bee is a worker bee that lays unfertilized eggs, usually in the absence of a queen bee. Only drones develop from the eggs of laying worker bees (with some exceptions, see thelytoky). A beehive cannot survive with only a laying worker bee. Prevalence Even in a normal hive, about 1% of workers have ovaries developed enough to lay eggs. However the usual number of the laid eggs is very small. Only eight eggs (seven moderately and one fully developed) were found after examining of 10,634 worker bees (strong colony contains about 100,000). Workers eventually lay significant numbers of eggs only in queenless colonies. Development Laying workers develop in the absence of open brood as produced by a healthy adult queen. Normally, pheromones from the brood – known as brood recognition pheromones – prevent development of the workers' ovaries. Laying workers can develop after the colony's queen has been lost to swarming Swarm behaviour, or swar ...
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Worker Bee
A worker bee is any female (eusocial) bee that lacks the full reproductive capacity of the colony's queen bee; under most circumstances, this is correlated to an increase in certain non-reproductive activities relative to a queen. While worker bees occur in all eusocial bee species, the term is rarely used (outside of scientific literature) for any bees other than honey bees. Honey bee workers gather pollen into the pollen baskets on their back legs and carry it back to the hive where it is used as food for the developing brood. Pollen carried on their bodies may be carried to another flower where a small portion can rub off onto the pistil, resulting in cross pollination. A significant amount of the world's food supply, particularly fruit, depends greatly on crop pollination by honey bees. Nectar is sucked up through the proboscis, mixed with enzymes in the stomach, and carried back to the hive, where it is stored in wax cells and evaporated into honey. Life cycle Although the ...
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Fertilization
Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Processes such as insemination or pollination which happen before the fusion of gametes are also sometimes informally called fertilisation. The cycle of fertilisation and development of new individuals is called sexual reproduction. During double fertilisation in angiosperms the haploid male gamete combines with two haploid polar nuclei to form a triploid primary endosperm nucleus by the process of vegetative fertilisation. History In Antiquity, Aristotle conceived the formation of new individuals through fusion of male and female fluids, with form and function emerging gradually, in a mode called by him as epigenetic. In 1784, Spallanzani established the need of interaction between the female's ovum and male's sperm to form a zygote in frog ...
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Queen Bee
A queen bee is typically an adult, mated female (gyne) that lives in a colony or hive of honey bees. With fully developed reproductive organs, the queen is usually the mother of most, if not all, of the bees in the beehive. Queens are developed from larvae selected by worker bees and specially fed in order to become sexually mature. There is normally only one adult, mated queen in a hive, in which case the bees will usually follow and fiercely protect her. The term "queen bee" can be more generally applied to any dominant reproductive female in a colony of a eusocial bee species other than honey bees. However, as in the Brazilian stingless bee ''Schwarziana quadripunctata'', a single nest may have multiple queens or even dwarf queens, ready to replace a dominant queen in a case of sudden death. Development During the warm parts of the year, female "worker" bees leave the hive every day to collect nectar and pollen. While male bees serve no architectural or pollinating purpo ...
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Drone (bee)
A drone is a male honey bee. Unlike the female worker bee, drones do not have stingers. They gather neither nectar nor pollen and are unable to feed without assistance from worker bees. A drone's only role is to mate with a maiden queen in nuptial flight. Genetics Drones carry only one type of allele at each chromosomal position, because they are haploid (containing only one set of chromosomes from the mother). During the development of eggs within a queen, a diploid cell with 32 chromosomes divides to generate haploid cells called gametes with 16 chromosomes. The result is a haploid egg, with chromosomes having a new combination of alleles at the various loci. This process is called arrhenotokous parthenogenesis or simply arrhenotoky. Because the male bee technically has only a mother, and no father, its genealogical tree is unusual. The first generation has one member (the male). One generation back also has one member (the mother). Two generations back are two members (the ...
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Egg (biology)
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the animal hatches. Most arthropods such as insects, vertebrates (excluding live-bearing mammals), and mollusks lay eggs, although some, such as scorpions, do not. Reptile eggs, bird eggs, and monotreme eggs are laid out of water and are surrounded by a protective shell, either flexible or inflexible. Eggs laid on land or in nests are usually kept within a warm and favorable temperature range while the embryo grows. When the embryo is adequately developed it hatches, i.e., breaks out of the egg's shell. Some embryos have a temporary egg tooth they use to crack, pip, or break the eggshell or covering. The largest recorded egg is from a whale shark and was in size. Whale shark eggs typically hatch within the mother. At and up to , the o ...
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Thelytoky
Thelytoky (from the Greek ''thēlys'' "female" and ''tokos'' "birth") is a type of parthenogenesis in which females are produced from unfertilized eggs, as for example in aphids. Thelytokous parthenogenesis is rare among animals and reported in about 1,500 species, about 1 in 1000 of described animal species, according to a 1984 study. It is more common in invertebrates, like arthropods, but it can occur in vertebrates, including salamanders, fish, and reptiles such as some whiptail lizards. Thelytoky can occur by different mechanisms, each of which has a different impact on the level of homozygosity. It is found in several groups of Hymenoptera, including Apidae, Aphelinidae, Cynipidae, Formicidae, Ichneumonidae, and Tenthredinidae. It can be induced in Hymenoptera by the bacteria ''Wolbachia'' and ''Cardinium''. Arrhenotoky and thelytoky in Hymenoptera Hymenoptera (ants, bees, wasps, and sawflies) have a haplodiploid sex-determination system. They produce haploid males ...
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Beehive (beekeeping)
A beehive is an enclosed structure in which some honey bee species of the subgenus '' Apis'' live and raise their young. Though the word ''beehive'' is commonly used to describe the nest of any bee colony, scientific and professional literature distinguishes ''nest'' from ''hive''. ''Nest'' is used to discuss colonies that house themselves in natural or artificial cavities or are hanging and exposed. ''Hive'' is used to describe an artificial/man-made structure to house a honey bee nest. Several species of ''Apis'' live in colonies, but for honey production the western honey bee (''Apis mellifera'') and the eastern honey bee (''Apis cerana'') are the main species kept in hives. The nest's internal structure is a densely packed group of hexagonal prismatic cells made of beeswax, called a honeycomb. The bees use the cells to store food (honey and pollen) and to house the brood (eggs, larvae, and pupae). Beehives serve several purposes: production of honey, pollination of near ...
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Pheromone
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavior of the receiving individuals. There are ''alarm signal, alarm pheromones'', ''food trail pheromones'', ''sex pheromones'', and many others that affect behavior or physiology. Pheromones are used by many organisms, from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been particularly well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones. The ecological functions and evolution of pheromones are a major topic of research in the field of chemical ecology. Background The portmanteau word "pheromone" was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek φερω ''pheroo'' ('I carry') and ὁρμων ''hormon'' ('stimulating'). P ...
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Pheromone (honeybee)
The pheromones of the honey bee are mixtures of chemical substances released by individual bees into the hive or environment that cause changes in the physiology and behaviour of other bees. Introduction Honey bees (''Apis mellifera'') have one of the most complex pheromonal communication systems found in nature, possessing 15 known glands that produce an array of compounds.Free, John B., ''Pheromones of social bees.'' Ithaca, N.Y.: Comstock, 1987.Blum, M.S. 1992. Honey bee pheromones in The Hive and the Honey Bee, revised edition (Dadant and Sons: Hamilton, Illinois), pages 385–389. These chemical messengers secreted by a queen, drone, worker bee or laying worker bee to elicit a response in other bees. The chemical messages are received by the bee's antenna and other body parts. They are produced as a volatile or non-volatile liquid and transmitted by direct contact as a liquid or vapor. Honey bee pheromones can be grouped into releaser pheromones which temporarily affect the ...
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Ovary
The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. The ovaries also secrete hormones that play a role in the menstrual cycle and fertility. The ovary progresses through many stages beginning in the prenatal period through menopause. It is also an endocrine gland because of the various hormones that it secretes. Structure The ovaries are considered the female gonads. Each ovary is whitish in color and located alongside the lateral wall of the uterus in a region called the ovarian fossa. The ovarian fossa is the region that is bounded by the external iliac artery and in front of the ureter and the internal iliac artery. This area is about 4 cm x 3 cm x 2 cm in size.Daftary, Shirish; Chakravarti, Sudip (2011). Manual of Obstetrics, 3rd Edition. Elsevier. pp. 1-16. . The ovari ...
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Swarming (honey Bee)
Swarming is a honey bee colony's natural means of reproduction. In the process of swarming, a single colony splits into two or more distinct colonies. Swarming is mainly a spring phenomenon, usually within a two- or three-week period depending on the locale, but occasional swarms can happen throughout the producing season. Secondary ''afterswarms'', or ''cast swarms'' may happen. Cast swarms are usually smaller and are accompanied by a virgin queen. Sometimes a beehive will swarm in succession until it is almost totally depleted of workers. One species of honey bee that participates in such swarming behavior is ''Apis cerana.'' The reproduction swarms of this species settle away from the natal nest for a few days and will then depart for a new nest site after getting information from scout bees. Scout bees search for suitable cavities in which to construct the swarm's home. Successful scouts will then come back and report the location of suitable nesting sites to the other bees. ...
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