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Kokartus
''Kokartus'' is an extinct genus of prehistoric stem-group salamander (Caudata) from the Middle Jurassic Balabansai Formation of Kyrgyzstan. The absence of clear lines of arrested growth and annuli in long bones suggests that the animals lived in an environment with stable local conditions. Like other members of Karauridae, it is thought to have been neotenic. ''Kokartus'' is thought to have fed using suction feeding via the enlargement of the buccal cavity on small fish and invertebrates, with the well developed palatal dentition (teeth on the roof of the mouth) and marginal teeth helping to grasp prey. See also * Prehistoric amphibian * List of prehistoric amphibians This list of prehistoric amphibians is an attempt to create a comprehensive listing of all genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accepted g ... References Prehistoric salamanders Middle Jurassic amp ...
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Karauridae
The Karauridae are a family of stem-group salamanders (Caudata) that are known from the Middle Jurassic to Late Jurassic in Central Asia (Kazakhstan and Kyrgyzstan) and Western Europe (United Kingdom). The family includes three members: ''Karaurus'' from the Middle-Late Jurassic Karabastau Formation of Kazakhstan, ''Kokartus'' from the Middle Jurassic Balabansai Formation of Kyrgyzstan, and ''Marmorerpeton'' from the Middle Jurassic Forest Marble Formation of England and Kilmaluag Formation of Scotland. The members are some of the oldest known salamanders. The family is united by several morphological characters, including sculptured skull roof bones. Like some modern salamanders, karaurids were neotenic. Members of the family likely fed via suction feeding Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This probl ...
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Salamander
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela. Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders rarely have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults. This group of amphibians is capable of regenerating lost lim ...
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Caudata
The Caudata are a group of amphibians containing the extant salamanders (Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. Evolution The origins and evolutionary relationships between the three main groups of amphibians ( apodans, urodeles and anurans) is a matter of debate. A 2005 molecular phylogeny, based on rDNA analysis, suggest ...
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Balabansai Formation
The Balabansai Formation (sometimes transcribed Balabansay, also known as Balabansai Svita) is a geological formation in Kazakhstan, Kyrgyzstan and Uzbekistan whose strata date back to the Bathonian and Callovian stages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Jurassic, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 550–552. . The lithology primarily consists of variegated sandstones, siltstones, claystones, and rare gravels and marls. Many taxa have been found in the formation, including amphibians and mammals Vertebrate paleofauna Indeterminate Avialae remains located in Ongtustik Qazaqstan Oblysy, Kazakhstan. Fish Amphibians Turtles Lizards Choristoderes Crocodyliformes Dinosaurs Pterosaurs Mammals Invertebrate ...
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Prehistoric Amphibian
This list of prehistoric amphibians is an attempt to create a comprehensive listing of all Genus, genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (''nomen dubium, nomina dubia''), or were not formally published (''nomen nudum, nomina nuda''), as well as synonym (zoology), junior synonyms of more established names, and genera that are no longer considered amphibians. Modern forms are excluded from this list. The list currently includes 454 names. Naming conventions and terminology Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include: * Synonym (zoology), Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type (zoology), type specimens are later assigned to th ...
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List Of Prehistoric Amphibians
This list of prehistoric amphibians is an attempt to create a comprehensive listing of all genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful ('' nomina dubia''), or were not formally published (''nomina nuda''), as well as junior synonyms of more established names, and genera that are no longer considered amphibians. Modern forms are excluded from this list. The list currently includes 454 names. Naming conventions and terminology Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include: * Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synon ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Kyrgyzstan
Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east. Its capital and largest city is Bishkek. Ethnic Kyrgyz make up the majority of the country's seven million people, followed by significant minorities of Uzbeks and Russians. The Kyrgyz language is closely related to other Turkic languages. Kyrgyzstan's history spans a variety of cultures and empires. Although geographically isolated by its highly mountainous terrain, Kyrgyzstan has been at the crossroads of several great civilizations as part of the Silk Road along with other commercial routes. Inhabited by a succession of tribes and clans, Kyrgyzstan has periodically fallen under larger domination. Turkic nomads, who trace their ancestry to many Turkic states. It was first established as the Yenisei Kyrgyz Khaganate later in the ...
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Neoteny
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compared to other primates. In progenesis or paedogenesis, sexual development is accelerated. Both neoteny and progenesis result in paedomorphism (as having the form typical of children) or paedomorphosis (changing towards forms typical of children), a type of heterochrony. It is the retention in adults of traits previously seen only in the young. Such retention is important in evolutionary biology, domestication and evolutionary developmental biology. Some authors define paedomorphism as the retention of larval traits, as seen in salamanders.Schell, S. C. ''Handbook of Trematodes of North America North of Mexico'', 1985, pg. 22 History and etymology The origins of the concept of neoteny have been traced to the Bible (as argued by Ashley Monta ...
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Suction Feeding
Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This problem was first identified by Robert McNeill Alexander. As a result, underwater predators, especially bony fish, have evolved a number of specialized feeding mechanisms, such as filter feeding, ram feeding, suction feeding, protrusion, and pivot feeding. Most underwater predators combine more than one of these basic principles. For example, a typical generalized predator, such as the cod, combines suction with some amount of protrusion and pivot feeding. Suction feeding Suction feeding is a method of ingesting a prey item in fluids by sucking the prey into the predator's mouth. It is a highly coordinated behavior achieved by the dorsal rotation of the dermatocranium, lateral expansion of the suspensorium, and the depression of the lower jaw and hyoid. Suct ...
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