Kodymirus
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Kodymirus
''Kodymirus'' is a problematic genus of Early Cambrian arthropod, known from the Czech Republic, which bears some resemblance to eurypterids and aglaspidids. Although it possessed great appendage-like raptorial arms, it was not homologous with those of megacheirans. It is part of a small and low-diversity Paseky Shale fauna group, which dwelt in brackish waters. Description ''Kodymirus'' was a small predator at long. Its distinctive feature is pair of large, serrated raptorial appendages. These appendages appear convergent to those of megacheirans and radiodonts, but are not homologous as they postantennular, suggesting raptorial arms evolved multiple times in Cambrian arthropods. ''Kodymirus'' is an enigmatic arthropod, and its precise taxonomic affinity remains uncertain. It was initially believed to be an early eurypterid and later a megacheiran, but today it is best considered to be a stem-group or ally of the aglaspidids. Its basipods closely resembles those of '' ...
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Artiopoda
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods, with the proximal lobe being elongate and bearing a lamella, while the distal lobe is paddle-shaped and setiforous (bearing hair-or bristle like structures). The limb endopod has se ...
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Chelicerata Incertae Sedis
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered i ...
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Radiodonta
Radiodonta is an extinct Order (biology), order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris, Anomalocaris canadensis'', ''Hurdia, Hurdia victoria'', ''Peytoia nathorsti'', ''Titanokorys gainesii, Titanokorys gainessii, Cambroraster, Cambroraster falcatus'' and ''Amplectobelua, Amplectobelua symbrachiata'', the Ordovician ''Aegirocassis, Aegiroca ...
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Fossils Of The Czech Republic
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Intertidal Zone
The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species of life, such as seastars, sea urchins, and many species of coral with regional differences in biodiversity. Sometimes it is referred to as the ''littoral zone'' or '' seashore'', although those can be defined as a wider region. The well-known area also includes steep rocky cliffs, sandy beaches, bogs or wetlands (e.g., vast mudflats). The area can be a narrow strip, as in Pacific islands that have only a narrow tidal range, or can include many meters of shoreline where shallow beach slopes interact with high tidal excursion. The peritidal zone is similar but somewhat wider, extending from above the highest tide level to below the lowest. Organisms in the intertidal zone are adapted to an environment of harsh extremes, living in water pr ...
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Estuary
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environments and are an example of an ecotone. Estuaries are subject both to marine influences such as tides, waves, and the influx of saline water, and to fluvial influences such as flows of freshwater and sediment. The mixing of seawater and freshwater provides high levels of nutrients both in the water column and in sediment, making estuaries among the most productive natural habitats in the world. Most existing estuaries formed during the Holocene epoch with the flooding of river-eroded or glacially scoured valleys when the sea level began to rise about 10,000–12,000 years ago. Estuaries are typically classified according to their geomorphological features or to water-circulation patterns. They can have many different names, such as bays, ...
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Beckwithia Typa
''Beckwithia'' is an extinct genus of average to large size ( to ), soft bodied artiopodan arthropods that lived during the late Middle and Upper Cambrian on the former paleocontinent Laurentia, in what are now Utah and Wisconsin. It has an inverted egg-shape, with a semi-circular head, 11 abdominal segments and a tail spine. It is a typical member of the group Aglaspidida. Etymology ''Beckwithia '' was named after Frank Beckwith, editor and publisher of the Millard County Chronicle of Delta, Utah in the early to middle 1900s, who had a passion for trilobites. Distribution ''Beckwithia'' has been collected from the Middle and Upper Cambrian of Utah (Weeks Formation, House Range, Millard County) and from the Franconian (in North-America called Sunwaptan) of Wisconsin. Taxonomic history ''Beckwithia'' was previously placed in a separate family, the Beckwithiidae, but it has now been established that the defining character (that the terminal segments appeared to be fused) i ...
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Emeraldella
''Emeraldella'' is a genus of arthropod known from the Middle Cambrian of North America. The type species ''E. brocki'' was described in 1912 from the Burgess Shale. 21 specimens of ''Emeraldella'' are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. A re-study on the species was done in 2012. A second species ''E. brutoni'' is known from the Wheeler Shale, which was described in 2011. An additional specimen of ''E. brutoni'' was described in 2019, which revealed more of the anatomy. It has been placed as part of the Artiopoda, a group of arthropods containing Trilobite, trilobites and their relatives.


Description

Caudal flaps are present on the terminal tergite, alongside an elongated spine.


''Emeraldella'' ''brocki''

''E. brocki'' is at least 60 mm in length, it possesses 12 tergites in t ...
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Aglaspidida
Aglaspidida is an extinct order of aquatic arthropods that were once regarded as primitive chelicerates. However, anatomical comparisons demonstrate that the aglaspidids cannot be accommodated within the chelicerates, and that they lie instead within the Artiopoda, thus placing them closer to the trilobites. Aglaspidida contains the subgroups Aglaspididae and Tremaglaspididae, which are distinguished by the presence of acute/spinose genal angles and a long spiniform tailspine in the Aglaspididae. Aglaspidid fossils are found in North America (upper Mississippi valley, Missouri and Utah), Europe, Australia, and China. Certain other artiopods are believed to be closely related to the aglaspidids, including the order Strabopida Strabopidae is the only family of the order Strabopida, a group of arachnomorph arthropods. The order Strabopida contains three valid genera and one doubtful, ''Khankaspis'', '' Paleomerus'', '' Parapaleomerus'' and '' Strabops'', all assigned in ..., w ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Incertae Sedis
' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by ' (of uncertain family), ' (of uncertain suborder), ' (of uncertain order) and similar terms. Examples *The fossil plant '' Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil ''Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Bocage's longbill, ''Motacilla bocagii' ...
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Cambrian Series 2
Cambrian Series 2 is the unnamed 2nd series of the Cambrian. It lies above the Terreneuvian series and below the Miaolingian. Series 2 has not been formally defined by the International Commission on Stratigraphy, lacking a precise lower boundary and subdivision into stages. The proposed lower boundary is the first appearance of trilobites which is estimated to be around million years ago. Naming The International Commission on Stratigraphy has not named the 2nd series of the Cambrian yet. In part the new name will replace the older terms "Lower Cambrian" and "Early Cambrian". The nomenclature used in Siberia uses the term "Yakutian" for this series. Subdivisions The 2nd series is currently subdivided by the ICS into two stages: Cambrian Stage 3 and Cambrian Stage 4. Both of these stages also lack formal definition. The Siberian nomenclature distinguishes three stages (lowest first): Atdabanian, Botomian and Toyonian. In general most subdivisions of this series rely on biostrati ...
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