Keuppia Levante
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Keuppia Levante
''Keuppia'' is an extinct genus of octopus. Taxonomy It consists of two species, ''Keuppia hyperbolaris'' and ''Keuppia levante'', both of which lived approximately 95 million years ago. Both species were found in fossilized form, which is very uncommon for extinct octopuses, as the soft tissue of dead octopuses almost always disintegrates before it has a chance to fossilize.Rare fossil octopuses found
NBC News, March 18, 2009 These fossils, along with those of the genus '' Styletoctopus'', were found from the -age Hâqel and

Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Octopus
An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like other cephalopods, an octopus is bilaterally symmetric with two eyes and a beaked mouth at the center point of the eight limbs. The soft body can radically alter its shape, enabling octopuses to squeeze through small gaps. They trail their eight appendages behind them as they swim. The siphon is used both for respiration and for locomotion, by expelling a jet of water. Octopuses have a complex nervous system and excellent sight, and are among the most intelligent and behaviourally diverse of all invertebrates. Octopuses inhabit various regions of the ocean, including coral reefs, pelagic waters, and the seabed; some live in the intertidal zone and others at abyssal depths. Most species grow quickly, mature ea ...
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Styletoctopus
''Styletoctopus'' is an extinct genus of octopus. The genus consists of the single species ''Styletoctopus annae'', which lived approximately 95 million years ago. Very few octopus species appear in the fossil record, as octopuses consist of soft tissue that usually decomposes before it has time to fossilize.Rare fossil octopuses found
NBC News, March 18, 2009


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2009 extinct fossil octopus discoveries Arthropods Cephalopods Three new species of extinct Octopoda discovered in 2009. The species – '' Keuppia hyperbolaris'', '' Keuppia levante'', and '' Styletoctopus annae'' – lived about 95 million years ago, and ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Hjoula
Hjoula ( ar, حجولا) is a municipality in the Byblos District of Keserwan-Jbeil Governorate, Lebanon. It is 70 kilometers north of Beirut. Hjoula has an elevation of between 920 and 1,100 meters above sea level. Hjoula has a total land area of 528 hectares. The village of Hjoula is known for its fertile soil and its woods, as well as Early Cretaceous fossils. Etymology The word is an Aramaic one, however the meaning is uncertain. Most historians and linguists suggest it means "Oval" due to its oval shape, while some others speculate it comes from the Syriac root "G-l-a" and subsequently to the word "Guola" which supposedly means "The place of the wandering salesman". History Many old relics and monuments were found in Hjoula. There was found a Phoenician inscription in "Jarabta". A Roman statue that represented a shepherd and his sheep was also found beside the lake, and was dated to the era of emperor Tiberius c. 20 CE; along small sets of coins that were written in Kufic wi ...
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Lebanon
Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus lies to its west across the Mediterranean Sea; its location at the crossroads of the Mediterranean Basin and the Arabian hinterland has contributed to its rich history and shaped a cultural identity of religious diversity. It is part of the Levant region of the Middle East. Lebanon is home to roughly six million people and covers an area of , making it the second smallest country in continental Asia. The official language of the state is Arabic, while French is also formally recognized; the Lebanese dialect of Arabic is used alongside Modern Standard Arabic throughout the country. The earliest evidence of civilization in Lebanon dates back over 7000 years, predating recorded history. Modern-day Lebanon was home to the Phoenicians, a m ...
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Gladius (cephalopod)
The gladius (plural: ''gladii''), or pen, is a hard internal bodypart found in many cephalopods of the superorder Decapodiformes (particularly squids) and in a single extant member of the Octopodiformes, the vampire squid (''Vampyroteuthis infernalis''). It is so named for its superficial resemblance to the Roman short sword of the same name, and is a vestige of the ancestral mollusc shell, which was external. The gladius is located dorsally within the mantle and usually extends for its entire length. Composed primarily of chitin, it lies within the shell sac, which is responsible for its secretion. Gladii are known from a number of extinct cephalopod groups, including teudopseids (''e.g.'' '' Actinosepia'', '' Glyphiteuthis'', '' Muensterella'', '' Palaeololigo'', '' Teudopsinia'', ''Teudopsis'', and ''Trachyteuthis''), loligosepiids (''e.g.'' '' Geopeltis'', '' Jeletzkyteuthis'', and '' Loligosepia''), and prototeuthids (''e.g.'' ''Dorateuthis'', '' Paraplesioteuthis'', ...
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2009 Extinct Fossil Octopus Discoveries
Arthropods Cephalopods Three new species of extinct Octopoda discovered in 2009. The species – '' Keuppia hyperbolaris'', '' Keuppia levante'', and '' Styletoctopus annae'' – lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the Octopoda order has remained relatively unchanged for tens of millions of years. The fossils included evidence of arms, muscles, rows of suckers, ink, and internal gills. The discovery was made by a team led by Dirk Fuchs of the Freie University, which is located at Berlin, Germany.Rare fossil octopuses found
NBC News, March 18, 2009
The fossils were found at Hakel and Hadjoula,

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Octopuses
An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like other cephalopods, an octopus is bilaterally symmetric with two eyes and a beaked mouth at the center point of the eight limbs. The soft body can radically alter its shape, enabling octopuses to squeeze through small gaps. They trail their eight appendages behind them as they swim. The siphon is used both for respiration and for locomotion, by expelling a jet of water. Octopuses have a complex nervous system and excellent sight, and are among the most intelligent and behaviourally diverse of all invertebrates. Octopuses inhabit various regions of the ocean, including coral reefs, pelagic waters, and the seabed; some live in the intertidal zone and others at abyssal depths. Most species grow quickly, mature early, a ...
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Prehistoric Cephalopod Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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