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Kalthifrons
''Kalthifrons'' is an extinct monospecific genus of mekosuchine crocodylian known from the Pliocene Tirari Formation in Australia. It contains a single species, ''Kalthifrons aurivellensis''. Discovery and naming The remains of ''Kalthifrons'' were discovered on the western shores of Lake Palankarinna in South Australia's Lake Eyre Basin. The holotype (specimen ''SAM P35062''), a cranium and two mandibular rami, was specifically recovered from the Mampuwordu Sand Member of the Pliocene Tirari Formation. It was found upside down with most of the ventral surface and both mandibular elements being in poor condition. The Golden Fleece Locality, where the fossil was discovered, also yielded a variety of isolated teeth, osteoderms and vertebrae remains, none of which however can be confidently assigned to ''Kalthifrons''. The generic name derives from the Dieri word for "spear" (kalthi) and fons, meaning forehead, a name chosen in reference to the animal's elongated frontal proces ...
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Mekosuchinae
Mekosuchinae is an extinct clade of crocodilians from the Cenozoic of Australasia. They first appear in the fossil record in the Eocene in Australia, and survived until the arrival of humans: in the Pleistocene in Australia and within the Holocene in the Pacific islands of Fiji, New Caledonia and Vanuatu. Mekosuchine crocodiles are a diverse group. One of the last species, ''Mekosuchus inexpectatus'' from Holocene New Caledonia, may have been arboreal. The early Miocene species '' Harpacochampsa camfieldensis'' may have resembled a false gharial. Another mekosuchine fossil, currently undescribed, has been found in Miocene deposits from New Zealand. One genus, ''Mekosuchus'', managed to spread to the islands of the Pacific; it is believed to have island-hopped across the Coral Sea, moving first to a now submerged island known as Greater Chesterfield Island, then New Caledonia and onwards. In the Pleistocene, ''Quinkana'' was one of the top terrestrial predators of the Australian ...
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Mekosuchine
Mekosuchinae is an extinct clade of Crocodilia, crocodilians from the Cenozoic of Australasia. They first appear in the fossil record in the Eocene in Australia, and survived until the arrival of humans: in the Pleistocene in Australia and within the Holocene in the Pacific islands of Fiji, New Caledonia and Vanuatu. Mekosuchine crocodiles are a diverse group. One of the last species, ''Mekosuchus, Mekosuchus inexpectatus'' from Holocene New Caledonia, may have been arboreal. The early Miocene species ''Harpacochampsa, Harpacochampsa camfieldensis'' may have resembled a false gharial. Another mekosuchine fossil, currently undescribed, has been found in Miocene deposits from New Zealand. One genus, ''Mekosuchus'', managed to spread to the islands of the Pacific; it is believed to have island-hopped across the Coral Sea, moving first to a now submerged island known as Chesterfield Islands, Greater Chesterfield Island, then New Caledonia and onwards. In the Pleistocene, ''Quinkana'' ...
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Volia
''Volia'' is an extinct monospecific genus of mekosuchine crocodylians from Fiji named in 2002.; ; 2002: An extinct Pleistocene endemic mekosuchine crocodylian from Fiji. ''Journal of vertebrate paleontology'', 22: 612–628. It was around long. Notwithstanding its comparatively small size, it was probably the apex predator of the Pleistocene ecosystems of Fiji. Fossils of ''Volia athollandersoni'', the type and currently only known species, have been found in the Voli-Voli and Wainibuku Caves of Viti Levu Island. The holotype is housed in the collection of the Museum of New Zealand Te Papa Tongarewa. ''V. athollandersoni'' and the other large reptiles of Fiji may have been exterminated by human hunting soon after Fiji was colonized by ancient Polynesians. Etyomology ''V. athollandersoni'' is named after the New Zealand archaeologist Atholl Anderson. Phylogeny A 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigr ...
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Trilophosuchus
''Trilophosuchus'' ("Triple Crest Crocodile") is an extinct genus of mekosuchine crocodilian from Australia. Unlike living crocodilians, it is hypothesized to have been terrestrial. ''Trilophosuchus'' was approximately in length. It had a short skull with three ridges on top and large eyes. Fossils have been found at Riversleigh in north-western Queensland, and are Miocene in age. Only a single species has been described, the type species ''T. rackhami''. Discovery ''Trilophosuchus'' is known from a posterior portion of the skull, QM F16856, the holotype specimen. Several other isolated bones of the skull were found. The material was collected from the Ringtail Site of the Gag Plateau in Riversleigh, one of Australia's most famous fossil localities. The skull was uncovered in 1985 during an excavation by the University of New South Wales. The deposit in which ''Trilophosuchus'' were found are Early Miocene in age, roughly 20 million years old. The deposit, known as System C, ...
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Australosuchus
''Australosuchus'' is an extinct monospecific genus of crocodylian belonging to the subfamily Mekosuchinae. The type and only known species ''Australosuchus clarkae'' lived during the Late Oligocene and the Early Miocene of southern Australia. The generic name ''Australosuchus'' means "Southern crocodile". It was described in 1991 from fossil material discovered at Lake Palankarinna in South Australia. A 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data established the inter-relationships within Crocodilia, which was expanded upon in 2021 by Hekkala ''et al.'' using paleogenomics by extracting DNA from the extinct ''Voay''. The below cladogram from the latest studies shows the placement of ''Australosuchus'' within Mekosuchinae Mekosuchinae is an extinct clade of crocodilians from the Cenozoic of Australasia. They first appear in the fossil record in the Eocene in Australia, and survive ...
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Baru
''Baru'' is an extinct genus of Australian mekosuchine crocodilian. It was semi-aquatic, around 4 m (13 ft) in length. Being semi-aquatic its habitat was around fresh pools of water in wet forests, ambushing their prey, much like modern species. The word ''Baru'' is Aboriginal and means "crocodile's ancestor". Fossils have been found in Australia at Riversleigh in north-western Queensland and Alcoota in the Northern Territory. Species There are currently three valid species within the genus ''Baru''. The type species ''B. darrowi'' is known from the Middle Miocene of the Northern Territory and is the largest reaching size of 4–5 m in length. It is named after English actor Paul Darrow. Two older species, ''B. huberi'' and ''B. wickeni'' are known from the Late Oligocene Etadunna Formation of Queensland. In 2009, a Baru skull had been found at the Alcoota fossil site about 200km (125 miles) from Alice Springs, in the Northern Territory (NT) of Australia. It was t ...
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Quinkana
''Quinkana'' is an extinct genus of mekosuchine crocodylians that lived in Australia from about 28 million to about 10,000 years ago. Most attributed specimens have been found in Queensland. It is speculated to have been one of the top predators of Pleistocene Australia, along with megalania, the giant monitor lizard, and the modern saltwater crocodile. The genus and type species, ''Q. fortirostrum'' was named by paleontologist Ralph E. Molnar in 1981. Other species in the genus are ''Q. timara'' (1994), ''Q. babarra'' (1996), and ''Q. meboldi'' (1997). The name ''Quinkana'' comes from the "Quinkans", a legendary folk spirit from Gugu-Yalanji mythology. ''Quinkana'' is thought to possess long legs and had ziphodont teeth (lateromedially compressed, recurved, and serrated). The genus is distinguishable by the combination of these ziphodont teeth and a broad snout. It also has a unique alveolar (tooth socket) structure and a short anterior palatine process (upper mouth bone). ''Q ...
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Mekosuchus
''Mekosuchus'' is a genus of extinct Australasian crocodiles within the subfamily Mekosuchinae. They are believed to have been made extinct by the arrival of humans on the South Pacific islands where they lived. The species of this genus were small in size, 2 m in maximum length, and terrestrial, making them the last surviving group of fully terrestrial crocodilians, leaving only semi-terrestrial species such as the Cuban crocodile and the dwarves ''Osteolaemus'' and ''Paleosuchus''. Fossils of related mekosuchines, such as ''Trilophosuchus'', have been found from Miocene Australia (the earliest known mekosuchine is the Eocene genus ''Kambara''), while ''Quinkana'' survived until the arrival of humans.Roberts, R.G., Flannery, T.F., Ayliffe, L.K., Yoshida, H., Olley J.M., Prideaux, G.J., Laslett, G.M., Baynes, A., Smith, M.A., Jones, R., Smith, B.L. (2001). New Ages for the Last Australian Megafauna: Continent-Wide Extinction About 46,000 Years Ago. ''Science'' 292 (5523): 1888†...
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Kambara
''Kambara'' is an extinct genus of mekosuchine crocodylian that lived during the Eocene epoch in Australia. Description At around 55 million years old, remains of ''Kambara'' are among the oldest Tertiary fossils found in Australia (although there are some recent Cretaceous fossils that are twice that age). ''Kambara'' is the oldest known mekosuchine. The genus name comes from an Aboriginal term meaning "crocodile". There are currently four species of ''Kambara'' described: the type species ''K. murgonensis'' (Willis & Molnar, 1993), ''K. implexidens'' (Salisbury & Willis, 1996), ''K. molnari'' (Holt et al., 2005), and ''K. taraina'' (Buchanan, 2009). All four species have a generalised crocodylian body plan, growing to sizes similar to the modern Saltwater Crocodile, ''Crocodylus porosus''. ''Kambara'' shows an interesting characteristic of having multiple bite patterns within the same genus. ''Kambara murgonensis'' has a near complete overbite, ''K. implexidens'' a more interl ...
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Paludirex
''Paludirex'' (meaning "swamp king") is an Extinction (biology), extinct genus of mekosuchinae, mekosuchine crocodylians from the Pliocene and Pleistocene of Australia. Remains of this animal have been found in the Australian Fossil Mammal Sites (Riversleigh), Riversleigh lagerstätte of northwestern Queensland. It was a medium-sized crocodile, estimated to grow to at least 4 metres in length. Discovery and naming In 1886 Charles Walter De Vis informally described ''Pallimnarchus pollens'' based on fragmentary cranomandibular and osteoderm material discovered around 1860 that was mineralised by apatite. While this marked the first fossil crocodile ever described from Australia, the name was merely coined "out of convenience". Regardless the name came to widespread use with a variety of specimens being referred in addition to the syntype specimens. In 1997 a second species was named by Paul Willis and Ralph E. Molnar, Ralph Molnar, ''Pallimnarchus gracilis''. Given the fragmentar ...
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Polytomy
An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree that contains any multifurcations can be described as a multifurcating tree. Soft polytomies vs. hard polytomies Two types of polytomies are recognised, soft and hard polytomies. Soft polytomies are the result of insufficient phylogenetic information: though the lineages diverged at different times – meaning that some of these lineages are closer relatives than others, and the available data does not allow recognition of this. Most polytomies are soft, meaning that they would be resolved into a typical tree of dichotomies if better data were available. In contrast, a hard polytomy represents a true divergence event of three or more lineages. Applications Interpretations for a polytomy depend on the individuals, that are represe ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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