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Isolecithal
Isolecithal (''Greek'' iso = equal, lekithos = yolk) refers to the even distribution of yolk in the cytoplasm of ova of mammals and other vertebrates, notably fishes of the families Petromyzontidae, Amiidae, and Lepisosteidae. Isolecithal cells have two equal hemispheres of yolk. However, during cellular development, normally under the influence of gravity, some of the yolk settles to the bottom of the egg, producing an uneven distribution of yolky hemispheres. Such uneven cells are known as telolecithal and are common where there is sufficient yolk mass. In the absence of a large concentration of yolk, four major cleavage types can be observed in isolecithal cells: radial holoblastic, spiral holoblastic, bilateral holoblastic, and rotational holoblastic cleavage. These holoblastic cleavage planes pass all the way through isolecithal zygotes during the process of cytokinesis. Coeloblastula is the next stage of development for eggs that undergo this radial cleavage. In mammals, becau ...
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Holoblastic
In embryology, cleavage is the division of cells in the early development of the embryo, following fertilization. The zygotes of many species undergo rapid cell cycles with no significant overall growth, producing a cluster of cells the same size as the original zygote. The different cells derived from cleavage are called blastomeres and form a compact mass called the morula. Cleavage ends with the formation of the blastula, or of the blastocyst in mammals. Depending mostly on the concentration of yolk in the egg, the cleavage can be holoblastic (total or entire cleavage) or meroblastic (partial cleavage). The pole of the egg with the highest concentration of yolk is referred to as the vegetal pole while the opposite is referred to as the animal pole. Cleavage differs from other forms of cell division in that it increases the number of cells and nuclear mass without increasing the cytoplasmic mass. This means that with each successive subdivision, there is roughly half the cytopl ...
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Yolk
Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example because they are laid in situations where the food supply is sufficient (such as in the body of the host of a parasitoid) or because the embryo develops in the parent's body, which supplies the food, usually through a placenta. Reproductive systems in which the mother's body supplies the embryo directly are said to be matrotrophic; those in which the embryo is supplied by yolk are said to be lecithotrophic. In many species, such as all birds, and most reptiles and insects, the yolk takes the form of a special storage organ constructed in the reproductive tract of the mother. In many other animals, especially very small species such as some fish and invertebrates, the yolk material is not in a special organ, but inside the egg cell. As sto ...
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Cytoplasm
In cell biology, the cytoplasm is all of the material within a eukaryotic cell, enclosed by the cell membrane, except for the cell nucleus. The material inside the nucleus and contained within the nuclear membrane is termed the nucleoplasm. The main components of the cytoplasm are cytosol (a gel-like substance), the organelles (the cell's internal sub-structures), and various cytoplasmic inclusions. The cytoplasm is about 80% water and is usually colorless. The submicroscopic ground cell substance or cytoplasmic matrix which remains after exclusion of the cell organelles and particles is groundplasm. It is the hyaloplasm of light microscopy, a highly complex, polyphasic system in which all resolvable cytoplasmic elements are suspended, including the larger organelles such as the ribosomes, mitochondria, the plant plastids, lipid droplets, and vacuoles. Most cellular activities take place within the cytoplasm, such as many metabolic pathways including glycolysis, and proces ...
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Ovum
The egg cell, or ovum (plural ova), is the female reproductive cell, or gamete, in most anisogamous organisms (organisms that reproduce sexually with a larger, female gamete and a smaller, male one). The term is used when the female gamete is not capable of movement (non-motile). If the male gamete (sperm) is capable of movement, the type of sexual reproduction is also classified as oogamous. A nonmotile female gamete formed in the oogonium of some algae, fungi, oomycetes, or bryophytes is an oosphere. When fertilized the oosphere becomes the oospore. When egg and sperm fuse during fertilisation, a diploid cell (the zygote) is formed, which rapidly grows into a new organism. History While the non-mammalian animal egg was obvious, the doctrine ''ex ovo omne vivum'' ("every living nimal comes froman egg"), associated with William Harvey (1578–1657), was a rejection of spontaneous generation and preformationism as well as a bold assumption that mammals also reproduced via ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Vertebrates
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Petromyzontidae
The northern lampreys (Petromyzontidae) are a family of lampreys. Northern lampreys have the highest number of chromosomes (164–174) among vertebrates. Genera * '' Caspiomyzon'' * '' Entosphenus'' * ''Eudontomyzon'' * ''Ichthyomyzon'' * ''Lampetra'' * ''Lethenteron'' * ''Petromyzon'' * ''Tetrapleurodon ''Tetrapleurodon'' is a genus of lampreys that are Endemism, endemic to the Lerma River, Lerma–Lake Chapala, Chapala basin in west–central Mexico. Both species are Threatened species, threatened. Species There are two recognized species in t ...'' References External links FishBase.org: Details for family Petromyzontidae – the Northern lampreys Fish described in 1827 Fish families Fish of Asia Fish of Europe Fish of North America Fish of the Atlantic Ocean Fish of the Pacific Ocean . Taxa named by Antoine Risso {{jawless-fish-stub ...
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Amiidae
The Amiidae are a family of basal ray-finned fishes. The bowfin is the only species to survive today, although additional species in all four subfamilies of Amiidae are known from Jurassic, Cretaceous, and Eocene fossils. Bowfins are now found throughout eastern North America, typically in slow-moving backwaters, canals, and ox-bow lakes. When the oxygen level is low (as often happens in still waters), the bowfin can rise to the surface and gulp air into its swim bladder, which is lined with blood vessels and can serve as a primitive lung. Amiidae is a monophyletic group that has numerous synapomorphic characters. Amiidae were widespread and particularly rich in species during the Eocene era. During this era, they appeared to be confined almost exclusively to fresh water. Taxonomy The family is divided into four subfamilies, with 11 genera described: *Amiidae **Subfamily Amiinae ***Genus ''Amia'' ***Genus †'' Cyclurus'' ***Genus †'' Pseudoamiatus'' **Subfamily †Amiopsin ...
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Lepisosteidae
Gars are members of the family Lepisosteidae, which are the only surviving members of the Ginglymodi, an ancient holosteian group of ray-finned fish, which first appeared during the Triassic, over 240 million years ago. Gars comprise seven living species of fish in two genera that inhabit fresh, brackish, and occasionally marine waters of eastern North America, Central America and Cuba in the Caribbean, though extinct members of the family were more widespread. Gars have elongated bodies that are heavily armored with ganoid scales, and fronted by similarly elongated jaws filled with long, sharp teeth. Gars are sometimes referred to as "garpike", but are not closely related to pike, which are in the fish family Esocidae. All of the gars are relatively large fish, but the alligator gar (''Atractosteus spatula'') is the largest; the alligator gar often grows to a length over and a weight over , and specimens of up to in length have been reported. Unusually, their vascularised sw ...
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Telolecithal
Telolecithal (Greek: ''τέλος'' (telos) = end, ''λέκιθος'' (lekithos) = yolk), refers to the uneven distribution of yolk in the cytoplasm of ova found in birds, reptiles, fish, and monotremes. The yolk is concentrated at one pole of the egg separate from the developing embryo. This type of egg undergoes discoidal meroblastic cleavage, where yolk is not incorporated in the cells during cell division. See also *Centrolecithal *Isolecithal Isolecithal (''Greek'' iso = equal, lekithos = yolk) refers to the even distribution of yolk in the cytoplasm of ova of mammals and other vertebrates, notably fishes of the families Petromyzontidae, Amiidae, and Lepisosteidae. Isolecithal cells have ... ReferencesEarly Development in FishEarly Development in Birds

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Zygote
A zygote (, ) is a eukaryotic cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individual organism. In multicellular organisms, the zygote is the earliest developmental stage. In humans and most other anisogamous organisms, a zygote is formed when an egg cell and sperm cell come together to create a new unique organism. In single-celled organisms, the zygote can divide asexually by mitosis to produce identical offspring. German zoologists Oscar and Richard Hertwig made some of the first discoveries on animal zygote formation in the late 19th century. Humans In human fertilization, a released ovum (a haploid secondary oocyte with replicate chromosome copies) and a haploid sperm cell (male gamete) combine to form a single diploid cell called the zygote. Once the single sperm fuses with the oocyte, the latter completes the division of the second ...
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Cytokinesis
Cytokinesis () is the part of the cell division process during which the cytoplasm of a single eukaryotic cell divides into two daughter cells. Cytoplasmic division begins during or after the late stages of nuclear division in mitosis and meiosis. During cytokinesis the spindle apparatus partitions and transports duplicated chromatids into the cytoplasm of the separating daughter cells. It thereby ensures that chromosome number and complement are maintained from one generation to the next and that, except in special cases, the daughter cells will be functional copies of the parent cell. After the completion of the telophase and cytokinesis, each daughter cell enters the interphase of the cell cycle. Particular functions demand various deviations from the process of symmetrical cytokinesis; for example in oogenesis in animals the ovum takes almost all the cytoplasm and organelles. This leaves very little for the resulting polar bodies, which in most species die without function, ...
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