Iranotherium
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Iranotherium
''Iranotherium'' ("Beast of Iran") was a genus of large elasmotheriine rhinoceros, comparable in size to a modern white rhinoceros, white rhino. It is known from the Late Miocene (Tortonian) of Maragheh, Maragha, Iran and the middle part of the Paleobiota of the Liushu Formation, Liushu formation of northwestern China. It was a precursor to the related ''Sinotherium'' and may have been ultimately outcompeted by its descendant. This species is most well known for showing unique sexual dimorphism among Rhinoceros, rhinos. Description ''Iranotherium'' is only known from Skull, cranial material, and rare postcranial elements. Skull and Horn The skull is long and dorsally concave, with a length of 775 mm. The Parietal bone, parietal crests are broad, the nasal bone is long and wide, with a shallow U-shaped nasal notch, the Orbital fenestra, orbit is prominent and has a high position close to the skull roof, the first half of the zygomatic arch is narrow. The skulls also show ...
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Paleobiota Of The Liushu Formation
The Liushu formation is a geological formation in the Gansu province of China that spans up to 100 m thick and is widely distributed within the Linxia Basin, with a Paleomagnetism, paleomagnetic age of between 11 and 6.4 Year, Mya (Late Miocene, Upper/Late Miocene). The formation is divided into three parts which represent three different ages, generally referred to as the "upper part" (youngest, at 6-7 Year, mya), the "middle part" (intermediate, at 9 to 7.6 mya. Also known as the Dashengou fauna, and the "lower part" (oldest). Mammals Ungulata Perissodactyla The Liushu formation displays a lineage of Elasmotheriine rhinoceros throughout its stratigraphy. ''Iranotherium'' is restricted to the middle part of the Geological formation, formation but remains become absent in the overlying layers, the only Elasmotheriine that can be found in the upper layers is ''Sinotherium''. Artiodactyla Carnivora Caniformia Feliformia Glires Proboscidea Birds Ref ...
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Sinotherium
''Sinotherium'' ("Chinese Beast") is a genus of single-horned elasmotheriine rhinoceros that lived from the late Miocene (Tortonian - Messinian) to Early Pliocene. It was ancestral to ''Elasmotherium,'' demonstrating a very important evolutionary transition from nasal-horned elasmotheriines to frontal-horned elasmotheriines. Its fossils have been found in the Karabulak Formation of Kazakhstan, lower jaw and teeth have been found in Mongolia, and a partial skull is known from the upper part of the Liushu Formation of western China. ''Sinotherium'' diverged from the ancestral genus, ''Iranotherium'', first found in Iran, during the early Pliocene. Some experts prefer to lump ''Sinotherium'', and ''Iranotherium'' into ''Elasmotherium''. Discovery, History and Taxonomy Species The type species of ''Sinotherium'' is ''S. lagrelii''. It is also known to have an additional species from the Zaisan depression of Kazakhstan called ''S. zaisanensis'', however, doubt has been raised o ...
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Elasmotheriine
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to plu ...
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Rhinoceros
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Rhinoceros
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Orbital Fenestra
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium (facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the frontal ...
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Zygomatic Arch
In anatomy, the zygomatic arch, or cheek bone, is a part of the skull formed by the zygomatic process of the temporal bone (a bone extending forward from the side of the skull, over the opening of the ear) and the temporal process of the zygomatic bone (the side of the cheekbone), the two being united by an oblique suture (the zygomaticotemporal suture); the tendon of the temporal muscle passes medial to (i.e. through the middle of) the arch, to gain insertion into the coronoid process of the mandible (jawbone). The jugal point is the point at the anterior (towards face) end of the upper border of the zygomatic arch where the masseteric and maxillary edges meet at an angle, and where it meets the process of the zygomatic bone. The arch is typical of '' Synapsida'' (“fused arch”), a clade of amniotes that includes mammals and their extinct relatives, such as ''Moschops'' and '' Dimetrodon''. Structure The zygomatic process of the temporal arises by two roots: * an ''anter ...
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Male
Male (symbol: ♂) is the sex of an organism that produces the gamete (sex cell) known as sperm, which fuses with the larger female gamete, or ovum, in the process of fertilization. A male organism cannot reproduce sexually without access to at least one ovum from a female, but some organisms can reproduce both sexually and asexually. Most male mammals, including male humans, have a Y chromosome, which codes for the production of larger amounts of testosterone to develop male reproductive organs. Not all species share a common sex-determination system. In most animals, including humans, sex is determined genetically; however, species such as ''Cymothoa exigua'' change sex depending on the number of females present in the vicinity. In humans, the word ''male'' can also be used to refer to gender in the social sense of gender role or gender identity. Overview The existence of separate sexes has evolved independently at different times and in different lineages, an example ...
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Zygomatic Bone
In the human skull, the zygomatic bone (from grc, ζῠγόν, zugón, yoke), also called cheekbone or malar bone, is a paired irregular bone which articulates with the maxilla, the temporal bone, the sphenoid bone and the frontal bone. It is situated at the upper and lateral part of the face and forms the prominence of the cheek, part of the lateral wall and floor of the orbit, and parts of the temporal fossa and the infratemporal fossa. It presents a malar and a temporal surface; four processes (the frontosphenoidal, orbital, maxillary, and temporal), and four borders. Etymology The term ''zygomatic'' derives from the Ancient Greek , ''zygoma'', meaning "yoke". The zygomatic bone is occasionally referred to as the zygoma, but this term may also refer to the zygomatic arch. Structure Surfaces The ''malar surface'' is convex and perforated near its center by a small aperture, the zygomaticofacial foramen, for the passage of the zygomaticofacial nerve and vessels; below ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Hypsodont
Hypsodont is a pattern of dentition with high-crowned teeth and enamel extending past the gum line, providing extra material for wear and tear. Some examples of animals with hypsodont dentition are cows and horses; all animals that feed on gritty, fibrous material. The opposite condition is called brachydont. Evolution Since the morphology of the hypsodont tooth is suited to a more abrasive diet, hypsodonty was thought to have evolved concurrently with the spread of grasslands. Grass contains phytoliths, silica-rich granules, which wear away dental tissue more quickly. Analysis has shown however, that the development of this morphology is out of sync with the flourishing of grasslands. Instead, the ingestion of grit and soil is hypothesized to be the primary driver of hypsodonty (the Grit, not grass hypothesis). Morphology Hypsodont dentition is characterized by: * high-crowned teeth * A rough, flattish occlusal surface adapted for crushing and grinding * Cementum both above ...
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Nasal Bone
The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Each has two surfaces and four borders. Structure The two nasal bones are joined at the midline internasal suture and make up the bridge of the nose. Surfaces The ''outer surface'' is concavo-convex from above downward, convex from side to side; it is covered by the procerus and nasalis muscles, and perforated about its center by a foramen, for the transmission of a small vein. The ''inner surface'' is concave from side to side, and is traversed from above downward, by a groove for the passage of a branch of the nasociliary nerve. Articulations The nasal articulates with four bones: two of the cranium, the frontal and ethmoid, and two of the face, the opposite nasal and the maxilla. Other animals In primitive bony fish and tetrapod ...
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