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Incisoscutum
''Incisoscutum'' is an extinct genus of arthrodire placoderm from the Early Frasnian Gogo Reef, from Late Devonian Australia. The genus contains two species ''I. ritchiei'', named after Alex Ritchie, a palaeoichthyologist and senior fellow of the Australian Museum, and ''I. sarahae'', named after Sarah Long, daughter of its discoverer and describer, John A. Long. The genus is important in the study of early vertebrates as well-preserved fossilized embryos have been found in female specimens and ossified pelvic claspers found in males. This shows that viviparity and internal fertilization was common amongst these primitive jawed vertebrates, which are outside the crown group Gnathostomata. In a study of fossil remains, comparison of the ontogeny of fourteen dermal plates from '' Compagopiscis croucheri'' and the more derived species ''Incisoscutum ritchiei'' suggested that lengthwise growth occurs earlier in the ontogeny than growth in width, and that dissociated allometric het ...
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Compagopiscis
''Compagopiscis'' is an extinct genus of placoderm known from the Gogo Formation. It lived in the Upper Devonian (Frasnian stage) of Western Australia. ''Compagopiscis'' is one of the earliest known vertebrates to have teeth. The genus is monotypic, with its only species being ''Compagopiscis croucheri''. Phylogeny ''Compagopiscis'' was originally classified as a member of the family Plourdosteidae. However, phylogenetic analysis later found Plourdosteidae to be an invalid grouping, and the family was dismissed. ''Torosteus'' is now considered to be a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ...
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Gogo Reef
The Gogo Formation in the Kimberley region of Western Australia is a Lagerstätte that exhibits exceptional preservation of a Devonian reef community. The formation is named after Gogo Station, a cattle station where outcrops appear and fossils are often collected from, as is nearby Fossil Downs Station. History The reef, which now stands up abruptly in the western Australian desert (as the Windjana Limestone), was first identified in 1940 by paleontologist Curt Teichert, who discovered the first fossil fish from the region. Sedimentology Unweathered sections of the Gogo Formation are made of siltstone, shale and calcarenite with numerous limestone concretions. These concretions are resistant to weathering, producing extensive nodule fields on the ground in areas where the surrounding rock has eroded away. The Gogo sediments represent deep, hypoxic seafloor deposits in the vicinity of a large tropical reef composed primarily of algae and stromatoporoids during the Frasni ...
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Gogo Formation
The Gogo Formation in the Kimberley region of Western Australia is a Lagerstätte that exhibits exceptional preservation of a Devonian reef community. The formation is named after Gogo Station, a cattle station where outcrops appear and fossils are often collected from, as is nearby Fossil Downs Station. History The reef, which now stands up abruptly in the western Australian desert (as the Windjana Limestone), was first identified in 1940 by paleontologist Curt Teichert, who discovered the first fossil fish from the region. Sedimentology Unweathered sections of the Gogo Formation are made of siltstone, shale and calcarenite with numerous limestone concretions. These concretions are resistant to weathering, producing extensive nodule fields on the ground in areas where the surrounding rock has eroded away. The Gogo sediments represent deep, hypoxic seafloor deposits in the vicinity of a large tropical reef composed primarily of algae and stromatoporoids during the Fras ...
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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus ...
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Materpiscis Attenboroughi
''Materpiscis'' (Latin for ''mother fish'') is a genus of ptyctodontid placoderm from the Late Devonian located at the Gogo Formation of Western Australia. Known from only one specimen, it is unique in having an unborn embryo present inside the mother, with remarkable preservation of a mineralised placental feeding structure (umbilical cord). This makes ''Materpiscis'' the oldest known vertebrate to show viviparity, or giving birth to live young. Discovery and naming The holotype was found in the Kimberley area of northern Western Australia by Lindsay Hatcher during the 2005 expedition to the Gogo led by John Long of Museum Victoria. Fossils from the Gogo Formation are preserved in limestone nodules, so dilute acetic acid is used to dissolve the surrounding limestone and reveal the fossil, often preserved in three dimensions with minimal distortion. The species was named ''Materpiscis attenboroughi'' in honour of David Attenborough who first drew attention to the significance o ...
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Late Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Internal Fertilization
Internal fertilization is the union of an egg and sperm cell during sexual reproduction inside the female body. Internal fertilization, unlike its counterpart, external fertilization, brings more control to the female with reproduction. For internal fertilization to happen there needs to be a method for the male to introduce the sperm into the female's reproductive tract. Most taxa that reproduce by internal fertilization are gonochoric. In mammals, reptiles, and certain other groups of animals, this is done by copulation, an intromittent organ being introduced into the vagina or cloaca. In most birds, the cloacal kiss is used, the two animals pressing their cloacas together while transferring sperm. Salamanders, spiders, some insects and some molluscs undertake internal fertilization by transferring a spermatophore, a bundle of sperm, from the male to the female. Following fertilization, the embryos are laid as eggs in oviparous organisms, or continue to develop inside the ...
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Worm Cast
A worm cast is a structure created by worms, typically on soils such as those on beaches that gives the appearance of multiple worms. They are also used to trace worm's location. References External links

Soil science {{zoology-stub ...
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Ossification
Ossification (also called osteogenesis or bone mineralization) in bone remodeling is the process of laying down new bone material by cells named osteoblasts. It is synonymous with bone tissue formation. There are two processes resulting in the formation of normal, healthy bone tissue: Intramembranous ossification is the direct laying down of bone into the primitive connective tissue ( mesenchyme), while endochondral ossification involves cartilage as a precursor. In fracture healing, endochondral osteogenesis is the most commonly occurring process, for example in fractures of long bones treated by plaster of Paris, whereas fractures treated by open reduction and internal fixation with metal plates, screws, pins, rods and nails may heal by intramembranous osteogenesis. Heterotopic ossification is a process resulting in the formation of bone tissue that is often atypical, at an extraskeletal location. Calcification is often confused with ossification. Calcification is sy ...
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Australoptyctodus Gardineri
The ptyctodontids ("folded-teeth") are placoderms of the order Ptyctodontida, containing the family Ptyctodontidae. With their big heads, big eyes, reduced armor and long bodies, the ptyctodontids bore a superficial resemblance to modern day chimaeras (Holocephali). Their armor was reduced to a pattern of small plates around the head and neck. Like the extinct and related acanthothoracids, and the living and unrelated holocephalians, most of the ptyctodontids are thought to have lived near the sea bottom and preyed on shellfish. On account of their radically reduced armor, some paleontologists have suggested that the Ptyctodontida were not actually placoderms, but actual holocephalians, some primitive group of elasmobranch fish, or even were the ancestors of the holocephalians, including the chimaeras. Thorough anatomical examinations of whole fossil specimens reveal that the profound similarities between these two groups are actually very superficial. The major differences ...
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Ptyctodont
The ptyctodontids ("folded-teeth") are placoderms of the order Ptyctodontida, containing the family Ptyctodontidae. With their big heads, big eyes, reduced armor and long bodies, the ptyctodontids bore a superficial resemblance to modern day chimaeras (Holocephali). Their armor was reduced to a pattern of small plates around the head and neck. Like the extinct and related acanthothoracids, and the living and unrelated holocephalians, most of the ptyctodontids are thought to have lived near the sea bottom and preyed on shellfish. On account of their radically reduced armor, some paleontologists have suggested that the Ptyctodontida were not actually placoderms, but actual holocephalians, some primitive group of elasmobranch fish, or even were the ancestors of the holocephalians, including the chimaeras. Thorough anatomical examinations of whole fossil specimens reveal that the profound similarities between these two groups are actually very superficial. The major differences ...
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Annual Review Of Earth And Planetary Sciences
''Annual Review of Earth and Planetary Sciences'' is an annual peer-reviewed scientific journal published by Annual Reviews, which broadly covers Earth and planetary sciences, including geology, atmospheric sciences, climate, geophysics, environmental science, geological hazards, geodynamics, planet formation, and solar system origins. The co-editors are Katherine H. Freeman ( Pennsylvania State University) and Raymond Jeanloz (University of California, Berkeley). As of 2022, ''Journal Citation Reports'' gives the journal a 2021 impact factor of 16.304. History The ''Annual Review of Earth and Planetary Sciences'' was first published in 1973 by the nonprofit publisher Annual Reviews. The goal of the editorial committee was to produce critical review articles that condensed a large volume of research into a final product usable by students, specialists, and non-specialists. In the late 1990s it began publishing materials electronically. Format changes in 2006 included a sim ...
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