Hymenophyllopsidaceae
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Hymenophyllopsidaceae
The Cyatheaceae are a family of ferns, the scaly tree ferns, one of eight families in the order Cyatheales in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may defined much more broadly (Cyatheaceae ''sensu lato'') as the only family in the Cyatheales, with the PPG I family treated as the subfamily Cyatheoideae. The narrower circumscription is used in this article. The family includes the world's tallest tree ferns, which reach heights up to 20 m. They are also very ancient plants, appearing in the fossil record in the late Jurassic, though the modern genera likely appeared in the Cenozoic. Cyatheaceae are the largest family of tree ferns, including about 640 species. Cyatheaceae and Dicksoniaceae, together with Metaxyaceae and Cibotiaceae, do not form a strongly supported monophyletic group and could be paraphyletic, but several individual subgroups are well supported as being monophyletic. Cyatheaceae are leptosporangiate ferns, the m ...
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Cyathea Glauca
''Alsophila glaucifolia'', synonym ''Cyathea glauca'', is a species of tree fern endemic to Réunion. Little is known about this species. Habitat and related species ''A. glaucifolia'' grows at higher altitudes () and it is one of three species of tree fern that are indigenous to Réunion island. * '' Alsophila celsa'' (syn. ''Cyathea excelsa'') which also occurs in Mauritius, grows at slightly lower altitudes (200-1700m). Like ''A. glaucifolia'', its leaves are tripinnate, but the new fronds of ''A. celsa'' are scaleless; those of ''A. glaucifolia'' have red-brown scales. * '' Alsophila borbonica'' (syn. ''Cyathea borbonica'') is the only species with bipinnate fronds. Image:Cyathea glauca fronds.jpg, The fronds are tripinnate (branching to three levels). Image:Cyathea glauca.JPG, Growing under '' Acacia heterophylla'' in the forest of Bélouve, Réunion Image:Image-Cyathea glauca insertion of fronds.JPG, Detail of trunk and insertion of fronds. New growth is ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Leaflet (botany)
A leaflet (occasionally called foliole) in botany is a leaf-like part of a compound leaf. Though it resembles an entire leaf, a leaflet is not borne on a main plant stem or branch, as a leaf is, but rather on a leaf, petiole or a branch of the leaf. Compound leaves are common in many plant families and they differ widely in morphology (biology), morphology. The two main classes of compound leaf morphology are Leaf shape, palmate and pinnate. For example, a ''Cannabis, hemp'' plant has palmate compound leaves, whereas some species of ''Acacia sensu lato, Acacia'' have pinnate leaves. The ultimate free division (or leaflet) of a compound leaf, or a pinnate subdivision of a multipinnate leaf is called a pinnule or pinnula. Image:Ветвь акации.jpg, Pinnate leaf of a Fabaceae, legume with 10 leaflets Image:Mimosa Pudica.gif, ''Mimosa pudica'' folding leaflets inward. See also * Compound leaf References

Plant morphology {{botany-stub ...
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Pinnate
Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in patterns of erosion or stream beds. The term derives from the Latin word ''pinna'' meaning "feather", "wing", or "fin". A similar concept is "pectination," which is a comb-like arrangement of parts (arising from one side of an axis only). Pinnation is commonly referred to in contrast to "palmation," in which the parts or structures radiate out from a common point. The terms "pinnation" and "pennation" are cognate, and although they are sometimes used distinctly, there is no consistent difference in the meaning or usage of the two words.Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928 Plants Botanically, pinnation is an arrangement of discr ...
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Vernation
Vernation (from ''vernal'' meaning ''spring'', since that is when leaves spring forth in temperate regions) is the formation of new leaves or fronds. In plant anatomy, it is the arrangement of leaves in a bud. In pine species, new leaves are short and encased in sheaths. Each leaf bundle consists of 2 to 5 needles. All the leaves on one section of branch grow in length together. In cabbage species, new leaves are folded over, each covered by the previous leaf. Circinate vernation Circinate vernation is the manner in which most fern fronds emerge. As the fern frond is formed, it is tightly curled so that the tender growing tip of the frond (and each subdivision of the frond) is protected within a coil. At this stage it is called a '' crozier'' (after the shepherd's crook) or ''fiddlehead'' (after the scrollwork at the top of a violin). As the lower parts of the frond expand and toughen up, they begin to photosynthesize, supporting the further growth and expansion of the frond. ...
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Marattiaceae
Marattiaceae is the only family of extant (living) ferns in the order Marattiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), Marattiales is the only order in the subclass Marattiidae. The family has six genera and about 110 species. Many are different in appearance from other ferns, having large fronds and fleshy rootstocks. Description The Marattiaceae diverged from other ferns very early in their evolutionary history and are quite different from many plants familiar to people in temperate zones. Many of them have massive, fleshy rootstocks and the largest known fronds of any fern. The Marattiaceae is one of two groups of ferns traditionally known as eusporangiate ferns, meaning that the sporangium is formed from a group of cells as opposed to a leptosporangium in which there is a single initial cell. The large fronds characteristic of the group are most readily found in the genus ''Angiopteris'', native to Australasia, Madagascar and Oceania. These fro ...
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Rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow horizontally. The rhizome also retains the ability to allow new shoots to grow upwards. A rhizome is the main stem of the plant that runs underground horizontally. A stolon is similar to a rhizome, but a stolon sprouts from an existing stem, has long internodes, and generates new shoots at the end, such as in the strawberry plant. In general, rhizomes have short internodes, send out roots from the bottom of the nodes, and generate new upward-growing shoots from the top of the nodes. A stem tuber is a thickened part of a rhizome or stolon that has been enlarged for use as a storage organ. In general, a tuber is high in starch, e.g. the potato, which is a modified stolon. The term "tuber" is often used imprecisely and is sometimes applied to ...
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Monilophyte
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Leptosporangiate Ferns
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, and the largest of these, being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Cibotiaceae
''Cibotium'' (from Greek , ''kibṓtion'', "little chest" or "box"), also known as manfern, is a genus of 11 species of tropical tree ferns. It is the only genus in family Cibotiaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may be treated as the subfamily Cibotioideae of a very broadly defined family Cyatheaceae, the family placement used for the genus in '' Plants of the World Online'' . Species , '' Plants of the World Online'' accepted the following species and hybrids: Some extinct species have also been placed in this genus: *†''Cibotium iwatense'' Ogura *†''Cibotium oregonense'' Barrington Distribution Species of the genus are distributed fairly narrowly in Hawaii (four species, plus a hybrid, collectively known as ''hāpuu''), Southeast Asia (five species), and the cloud forests of Central America and Mexico (two species). The natural habitat of ''Cibotium'' is among the dripping trees and stream gullies of the ra ...
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Metaxyaceae
''Metaxya'' is a neotropical genus of ferns in the order Cyatheales. It is the only genus in the family Metaxyaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed in the subfamily Metaxyoideae of a more broadly defined family Cyatheaceae, the family placement used in ''Plants of the World Online'' . The species of the genus are characterized by large fronds that approach 8 ft (2.5 m) in length. Species , ''Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...'' accepted the following species: References External links * * Cyatheales Fern genera {{Cyatheales-stub ...
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Dicksoniaceae
Dicksoniaceae is a group of tropical, subtropical and warm temperate ferns, treated as a family in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), and counting 30-40 species. Alternatively, the family may be sunk into a very broadly defined family Cyatheaceae ''sensu lato'' as the subfamily Dicksonioideae. Most of the genera in the family are terrestrial ferns or have very short trunks compared to tree ferns of the family Cyatheaceae ''sensu stricto''. However, some of the larger species can reach several metres in height. A number of others are epiphytes. They are found mostly in tropical regions in the Southern Hemisphere, as far south as southern New Zealand. Larger tree ferns in the genus ''Cibotium'' were formerly included in Dicksoniaceae, but are now segregated as the family Cibotiaceae. Description Species in the family are generally characterized by large pinnate fronds, 1–4 m long. The family includes several species of tree ferns, which grow a single ...
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