Hicklingia
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Hicklingia
''Hicklingia'' is a genus of extinct plants of the Middle Devonian (around ). Compressed specimens were first described in 1923 from the Old Red Sandstone of Scotland., cited in Initially the genus was placed in the "rhyniophytes", but this group is defined as having terminal sporangia (spore-forming organs), and later work showed that the sporangia of ''Hicklingia'' were lateral rather than strictly terminal, so that it is now regarded as having affinities with the zosterophylls. Description The sporophyte had a tufted growth habit, with narrow leafless stems (axes) up to 17 cm high which branched dichotomously. Sporangia were borne on short stalks (up to 3 mm), on all sides of the stem and also terminally. There are oval scars on specimens where the stalks are presumed to have broken off. The lateral sporangia were closely adpressed to the stem. The effect is of a 'spike' of sporangia which terminates some stems. The sporangia opened via slits, but these did not ha ...
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Zosterophyll
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Zosterophylls
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Lycopodiophyta
The lycophytes, when broadly circumscribed, are a vascular plant (tracheophyte) subgroup of the kingdom Plantae. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineages of extant (living) vascular plants; the group contains extinct plants that have been dated from the Silurian (ca. 425 million years ago). Lycophytes were some of the dominating plant species of the Carboniferous period, and included tree-like species, although extant lycophytes are relatively small plants. The scientific names and the informal English names used for this group of plants are ambiguous. For example, "Lycopodiophyta" and the shorter "Lycophyta" as well as the informal "lycophyte" may be used to include the extinct zosterophylls or to exclude them. Description Lycophytes reproduce by spores and have alternation of generations in which (like other vascular plants) the sporophyte generation is dominant. Some lycophyte ...
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Lycophyte
The lycophytes, when broadly circumscribed, are a vascular plant (tracheophyte) subgroup of the kingdom Plantae. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineages of extant (living) vascular plants; the group contains extinct plants that have been dated from the Silurian (ca. 425 million years ago). Lycophytes were some of the dominating plant species of the Carboniferous period, and included tree-like species, although extant lycophytes are relatively small plants. The scientific names and the informal English names used for this group of plants are ambiguous. For example, "Lycopodiophyta" and the shorter "Lycophyta" as well as the informal "lycophyte" may be used to include the extinct zosterophylls or to exclude them. Description Lycophytes reproduce by spores and have alternation of generations in which (like other vascular plants) the sporophyte generation is dominant. Some lycophyte ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Middle Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Old Red Sandstone
The Old Red Sandstone is an assemblage of rocks in the North Atlantic region largely of Devonian age. It extends in the east across Great Britain, Ireland and Norway, and in the west along the northeastern seaboard of North America. It also extends northwards into Greenland and Svalbard. These areas were a part of the ancient continent of Euramerica, Euramerica/Laurussia. In Britain it is a lithostratigraphy, lithostratigraphic unit (a sequence of rock strata) to which Stratigraphy, stratigraphers accord Geological unit#Lithostratigraphic units, supergroup status and which is of considerable importance to early paleontology. For convenience the short version of the term, ORS is often used in literature on the subject. The term was coined to distinguish the sequence from the younger New Red Sandstone which also occurs widely throughout Britain. Sedimentology The Old Red Sandstone describes a suite of sedimentary rocks deposited in a variety of environments during the Devonian ...
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Scotland
Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the north and west, the North Sea to the northeast and east, and the Irish Sea to the south. It also contains more than 790 islands, principally in the archipelagos of the Hebrides and the Northern Isles. Most of the population, including the capital Edinburgh, is concentrated in the Central Belt—the plain between the Scottish Highlands and the Southern Uplands—in the Scottish Lowlands. Scotland is divided into 32 administrative subdivisions or local authorities, known as council areas. Glasgow City is the largest council area in terms of population, with Highland being the largest in terms of area. Limited self-governing power, covering matters such as education, social services and roads and transportation, is devolved from the Scott ...
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Rhyniophyte
The rhyniophytes are a group of extinct early vascular plants that are considered to be similar to the genus ''Rhynia'', found in the Early Devonian (around ). Sources vary in the name and rank used for this group, some treating it as the class Rhyniopsida, others as the subdivision Rhyniophytina or the division Rhyniophyta. The first definition of the group, under the name Rhyniophytina, was by Banks, since when there have been many redefinitions, including by Banks himself. "As a result, the Rhyniophytina have slowly dissolved into a heterogeneous collection of plants ... the group contains only one species on which all authors agree: the type species ''Rhynia gwynne-vaughanii''". When defined very broadly, the group consists of plants with dichotomously branched, naked aerial axes ("stems") with terminal spore-bearing structures (sporangia). The rhyniophytes are considered to be stem group tracheophytes (vascular plants). Definitions The group was described as a subdivision of ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Sporophyte
A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote produced when a haploid egg cell is fertilized by a haploid sperm and each sporophyte cell therefore has a double set of chromosomes, one set from each parent. All land plants, and most multicellular algae, have life cycles in which a multicellular diploid sporophyte phase alternates with a multicellular haploid gametophyte phase. In the seed plants, the largest groups of which are the gymnosperms and flowering plants (angiosperms), the sporophyte phase is more prominent than the gametophyte, and is the familiar green plant with its roots, stem, leaves and cones or flowers. In flowering plants the gametophytes are very reduced in size, and are represented by the germinated pollen and the embryo sac. The sporophyte produces spores (hence t ...
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Vascular Tissue
Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There are also two meristems associated with vascular tissue: the vascular cambium and the cork cambium. All the vascular tissues within a particular plant together constitute the vascular tissue system of that plant. The cells in vascular tissue are typically long and slender. Since the xylem and phloem function in the conduction of water, minerals, and nutrients throughout the plant, it is not surprising that their form should be similar to pipes. The individual cells of phloem are connected end-to-end, just as the sections of a pipe might be. As the plant grows, new vascular tissue differentiates in the growing tips of the plant. The new tissue is aligned with existing vascular tissue, maintaining its connection throughout the plant. The ...
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