Hamipteridae
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Hamipteridae
Hamipteridae (or hamipterids) is a small family of anhanguerian pterosaurs known from the Early Cretaceous of China and Spain. Classification The cladogram below follows the topology recovered by Pêgas ''et al.'' (2019). In the analysis, they assigned Hamipteridae as the sister taxon of the family Anhangueridae, both within the larger clade Anhangueria Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ....Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, References Pteranodontoids Prehistoric reptile families {{pterosaur-stub ...
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Anhangueria
Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Anhangueria was named by paleontologists Taissa Rodrigues and Alexander Kellner in a review of ''Ornithocheirus'' species in 2013, they defined the clade as a branch-based taxon consisting of all pteranodontoids more closely related to ''Anhanguera blittersdorffi'' than to ''Istiodactylus latidens'' and ''Cimoliopterus cuvieri''. Classification Anhangueria originally only contained the genera '' Brasileodactylus'', ''Camposipterus'', ''Cearadactylus'', ''Ludodactylus'' as well as the family Anhangueridae, however, recent analyses had recovered the family Hamipteridae within this clade as well. In 2014, paleontologist Brian Andres and colleagues assigned more groups and genera within this clade, this included ''Guidraco'', the subfa ...
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Anhangueria
Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Anhangueria was named by paleontologists Taissa Rodrigues and Alexander Kellner in a review of ''Ornithocheirus'' species in 2013, they defined the clade as a branch-based taxon consisting of all pteranodontoids more closely related to ''Anhanguera blittersdorffi'' than to ''Istiodactylus latidens'' and ''Cimoliopterus cuvieri''. Classification Anhangueria originally only contained the genera '' Brasileodactylus'', ''Camposipterus'', ''Cearadactylus'', ''Ludodactylus'' as well as the family Anhangueridae, however, recent analyses had recovered the family Hamipteridae within this clade as well. In 2014, paleontologist Brian Andres and colleagues assigned more groups and genera within this clade, this included ''Guidraco'', the subfa ...
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Iberodactylus
''Iberodactylus'' is a genus of pterodactyloid pterosaurs belonging to the clade Anhangueria, that during the Early Cretaceous lived in the area of present Spain. The type species is ''Iberodactylus andreui''. History of discovery and naming In the late 1980s, amateur paleontologist Javier Andreu discovered a pterosaur skull at the Los Quiñones site, west of Obón in Aragón. At the time it represented the most complete discovery of pterosaur fossil material in Spain apart from the remains of '' Prejanopterus''. In 2014, the find was reported in the scientific literature by José Antonio Ulloa-Rivas and identified as a member of the Ornithocheiroidea.Ulloa-Rivas, J.A. & Canudo, J.I. 2014. "New cranial remains of Ornithocheiroidea (Pterosauria) from the Barremian (Lower Cretaceous) of the Iberian Peninsula (Nuevos restos craneales de Ornithocheiroidea (Pterosauria) del Barremiense (Cretácico Inferior) de la Península Ibérica)". In: (Rodríguez, G.J.A. et alii, eds) ''New In ...
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Hamipterus Tianshanensis
''Hamipterus'' is an extinct genus of pteranodontoid pterosaurs from the Early Cretaceous Shengjinkou Formation of northwestern China. It is known from a single species, the type species, ''H. tianshanensis''. * Discovery and naming In 2006 from the Hami region in Xinjiang, the Shengjinkou Formation, a Konservat-Lagerstätte was reported, in this case lake sediments allowing for an exceptional preservation of fossils. The same year, Qiu Zhanxiang and Wang Banyue started official excavations. Part of the finds consisted of dense concentrations of pterosaur bones, associated with soft tissues and eggs. The site represented a nesting colony that storm floods had covered with mud. Dozens of individuals could be secured from a total that in 2014 was estimated to run into the many hundreds. In 2014, the type species ''Hamipterus tianshanensis'' was named and described by Wang Xiaolin, Alexander Kellner, Jiang Shunxing, Wang Qiang, Ma Yingxia, Yahefujiang Paidoula, Cheng Xin, Taissa Rod ...
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Hamipterus
''Hamipterus'' is an extinct genus of pteranodontoid pterosaurs from the Early Cretaceous Shengjinkou Formation of northwestern China. It is known from a single species, the type species, ''H. tianshanensis''. * Discovery and naming In 2006 from the Hami region in Xinjiang, the Shengjinkou Formation, a Konservat-Lagerstätte was reported, in this case lake sediments allowing for an exceptional preservation of fossils. The same year, Qiu Zhanxiang and Wang Banyue started official excavations. Part of the finds consisted of dense concentrations of pterosaur bones, associated with soft tissues and eggs. The site represented a nesting colony that storm floods had covered with mud. Dozens of individuals could be secured from a total that in 2014 was estimated to run into the many hundreds. In 2014, the type species ''Hamipterus tianshanensis'' was named and described by Wang Xiaolin, Alexander Kellner, Jiang Shunxing, Wang Qiang, Ma Yingxia, Yahefujiang Paidoula, Cheng Xin, Taissa R ...
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Caulkicephalus
''Caulkicephalus'' is a genus of anhanguerid pterosaur from the Isle of Wight off the coast of England. It lived during the Early Cretaceous period, about 130 million years ago. Discovery and naming Between 1995 and 2003, bone fragments of an unknown pterosaur were found at the Yaverland locality near Sandown. The discoveries were made in or from a brown clay layer from the Wessex Formation of the Wealden Group, stemming from the Early Cretaceous (Barremian age, about 130 million years ago). In 2005 a new genus was named and described by Lorna Steel, David Martill, David Unwin and John Winch. The type species is ''Caulkicephalus trimicrodon''. The genus name is a translation of "Caulkhead", a traditional nickname for Isle of Wight residents, partially derived from Greek ''kephale'', "head". The specific name, ''trimicrodon'', means "three small teeth", in reference to the dentition. Description The holotype is IWCMS 2002.189.1, 2, 4: three pieces, more or less contiguou ...
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Coloborhynchinae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (201 ...
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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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Anhanguerinae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (201 ...
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Ludodactylus
''Ludodactylus'' (meaning "play finger") is a genus of anhanguerid pterodactyloid pterosaur from the Early Cretaceous period (Aptian stage) of what is now the Crato Formation of the Araripe Basin in Ceará, Brazil. The type and only species is ''L. sibbicki''. The generic name ''Ludodactylus'' refers to the fact that the animal had the combination of teeth and a ''Pteranodon''-like head crest, similar to many toy pterosaurs, and no such creature was known to exist until the discovery of ''Ludodactylus''. However, ''Ludodactylus'' is not the only pterosaur known to possess these features, its very close relative ''Caulkicephalus'' is another example. Etymology The genus was named by Eberhard Frey ''et al.'' in 2003 and contains one known species, ''Ludodactylus sibbicki''. The name is derived from Latin ''ludus'', "play" and Greek ''daktylos'', "finger". ''Ludus'' refers to the fact, long lamented by paleontologists, that many toy pterosaurs combined teeth with a ''Pteranodon''-l ...
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Guidraco
''Guidraco'' (Chin. ''gui'' (鬼) "malicious ghost" + Lat. ''draco'' "dragon") is an extinct genus of toothed pterodactyloid pterosaur known from the Early Cretaceous of Liaoning Province, northeast China. According to many recent studies, ''Guidraco'' is a member of the group Anhanguerinae, a subfamily belonging to the larger group Anhangueridae. Discovery ''Guidraco'' is known only from the holotype IVPP V17083, an articulated partial skeleton consisting of a nearly complete skull, lower jaws and a series of four, second to fifth, cervical vertebrae. It was collected at Sihedang near Lingyuan City in the Liaoning Province from the Jiufotang Formation, dating to the Aptian stage of the Early Cretaceous, about 120 million years ago. Etymology ''Guidraco'' was first described and named by Wang Xiaolin, Alexander W.A. Kellner, Jiang Shunxing and Cheng Xin in 2012 and the type species is ''Guidraco venator''. The generic name is derived from Chinese ''gui'' (鬼), "m ...
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Uktenadactylus
''Uktenadactylus'' is a genus of anhanguerid pterodactyloid pterosaurs from the Lower Cretaceous Paw Paw Formation of Texas, United States and the Wessex Formation on the Isle of Wight, England. Fossil remains of ''Uktenadactylus'' dated back to the Early Cretaceous period (Barremian to Cenomanian stages), from about 125 to 100 million years ago. Discovery and naming In 1994, Yuong Nam-Lee named a new species within the genus ''Coloborhynchus'': ''Coloborhynchus wadleighi'', based on a partial snout found in 1992 in Albian layers in Tarrant County, holotype SMU 73058 ( Shuler Museum of Paleontology, Southern Methodist University at Dallas). The specific name honors the collector of the fossil, Chris Wadleigh. The reference of the species to the genus ''Coloborhynchus'' was based on the fact that both ''C. wadleighi'' and the type species of ''Coloborhynchus'', ''Coloborhynchus clavirostris'', share the trait of having three pairs of teeth laterally placed within a broad snout ti ...
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