Hyolithid
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Hyolithid
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs. Morphology The shell of a hyolith is typically one to four centimeters in length, triangular or elliptical in cross section. Some species have rings or stripes. It comprises two parts: the main conical shell (previously referred to as a ‘conch’) and a cap-like operculum. Some also had two curved supports known as ''helens'' They are calcareous – probably aragonitic All of these structures grew by marginal accretion. Shell microstructure The orthothecid shell has an internal layer with a microstructure of transverse bundles, and an external layer comprising longitudinal bundles. Helens Some hyoliths had helens, long structures that taper as they coil gently in a logarithmic spir ...
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Hyolithida
The Hyolithida are lophophorate The Lophophorata are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa. They have a lophophore. Molecular phylogenetic analyses suggest that lophophorates are protostomes Protostomia () is the clade of animals once though ...s, one of the two orders of hyolithid, the other being the Orthothecida. Most of our knowledge of the hyolithids comes from studies on the Hyolithida. Both orders had an operculum that was not hinged to the conch. However, the Hyolithida are distinct from the Orthothecida in having additional paired, curved, whiskerlike appendages. The Hyolithida were probably bottom feeders living in shallow water, and had tentacules. References Further reading * * Paleozoic invertebrates Prehistoric animal orders Hyolitha {{paleo-protostome-stub ...
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Hyoliths02
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs. Morphology The shell of a hyolith is typically one to four centimeters in length, triangular or elliptical in cross section. Some species have rings or stripes. It comprises two parts: the main conical shell (previously referred to as a ‘conch’) and a cap-like operculum. Some also had two curved supports known as ''helens'' They are calcareous – probably aragonitic All of these structures grew by marginal accretion. Shell microstructure The orthothecid shell has an internal layer with a microstructure of transverse bundles, and an external layer comprising longitudinal bundles. Helens Some hyoliths had helens, long structures that taper as they coil gently in a logarithmic spira ...
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Haplophrentis Burgess Shale
''Haplophrentis'' is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called ''helens'' protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts. Morphology Shell length of ''H. reesi'' reached up to while ''H. carinatus'' reached up to . Juveniles could of course be smaller. It is distinguished from '' Hyolithes'' by the presence of a longitudinal septum on the middle of the inner surface of the top of the shell. Its soft anatomy comprises 12(H. carinatus) to 16 (H. reesi) tentacles attached to a horseshoe-shaped lophophore. A pair of wide structures of uncertain function extend along the length of the conical shell. A larval shell is attached to the shell apex. Affinity The soft anatomy of ''Haplophrentis'' was key to establishing the hyoliths as members of the Lophophorata, the group ...
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Haplophrentis
''Haplophrentis'' is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called ''helens'' protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts. Morphology Shell length of ''H. reesi'' reached up to while ''H. carinatus'' reached up to . Juveniles could of course be smaller. It is distinguished from ''Hyolithes'' by the presence of a longitudinal septum on the middle of the inner surface of the top of the shell. Its soft anatomy comprises 12(H. carinatus) to 16 (H. reesi) tentacles attached to a horseshoe-shaped lophophore. A pair of wide structures of uncertain function extend along the length of the conical shell. A larval shell is attached to the shell apex. Affinity The soft anatomy of ''Haplophrentis'' was key to establishing the hyoliths as members of the Lophophorata, the group ...
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Mollusca
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine biology, marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater mollusc, freshwater and Terrestrial molluscs, terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8 Taxonomy (biology), taxonomic class (biology), classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurobiology, neurologically advanced of all inve ...
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Orthothecida
The orthothecids are one of the two hyolith Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as b ... orders. Marek diagnoses the order thus: Conchs with a flat or concave ventral surface — opercula with large, flat cardinal processes but without clavicles – tightly sigmoidal, sediment-filled intestine – helens absent. Sometimes the Circothecidae and Tetrathecidae are split out into a separate order 'Circothecida', which is defined by the bottom surface not being flat, the cardinal processes being pronounced, and a circular rim sometimes showing hints of differentiation into clavicles. Internal taxonomy Marek gives the following diagnoses: * Orthothecidae: Flat or concave bottom surface of conical shell. Transverse aperture. Kouchinsky lists the following taxonomic criteria: ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wife, establishing a quarry on the flanks of Fossil Ridge. The significance of soft-bodied preservation, and the range of organisms he recognised as new to science, led him to return to the quarry almost every year until 1924. At that point, aged 74, he had amassed over 65,000 ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups wer ...
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Lophophorata
The Lophophorata are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa. They have a lophophore. Molecular phylogenetic analyses suggest that lophophorates are protostomes, but on morphological grounds they have been assessed as deuterostome Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some ...s. Fossil finds of a segmented worm named Wufengella suggest that they evolved from a worm close to annelids. References {{Protostome-stub ...
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Sipunculid
The Sipuncula or Sipunculida (common names sipunculid worms or peanut worms) is a class containing about 162 species of unsegmented marine annelid worms. The name ''Sipuncula'' is from the genus name '' Sipunculus'', and comes from the Latin ''siphunculus'' meaning a "small tube". Sipuncula was once considered a phylum, but was demoted to a class of Annelida, based on recent molecular work. Sipunculans vary in size but most species are under in length. The body is divided into an unsegmented, bulbous trunk and a narrower, anterior section, called the "introvert", which can be retracted into the trunk. The mouth is at the tip of the introvert and is surrounded in most groups by a ring of short tentacles. With no hard parts, the body is flexible and mobile. Although found in a range of habitats throughout the world's oceans, the majority of species live in shallow water habitats, burrowing under the surface of sandy and muddy substrates. Others live under stones, in rock crev ...
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