Hughmilleriidae
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Hughmilleriidae
Hughmilleriidae (the name deriving from the type genus '' Hughmilleria'', which is named in honor of Scottish geologist Hugh Miller) is a family of eurypterids, an extinct group of aquatic arthropods. The hughmilleriids were the most basal members of the superfamily Pterygotioidea, in contrast with the more derived (more "advanced") families Pterygotidae and Slimonidae. Despite their classification as pterygotioids, the hughmilleriids possessed several characteristics shared with other eurypterid groups, such as the lanceolate telson (the most posterior segment of the body). Hughmilleriids are defined as pterygotioid eurypterids with swimming legs similar to those of the type genus, ''Hughmilleria'' (that is, 7th and 8th leg segments narrow and 9th segment very small), and whose second to fifth pair of appendages were spiniferous. Some further diagnostic characters unite the group, such as the slightly enlarged chelicerae (frontal appendages) and the streamlined shape of their b ...
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Hughmilleriidae Size
Hughmilleriidae (the name deriving from the type genus '' Hughmilleria'', which is named in honor of Scottish geologist Hugh Miller) is a family of eurypterids, an extinct group of aquatic arthropods. The hughmilleriids were the most basal members of the superfamily Pterygotioidea, in contrast with the more derived (more "advanced") families Pterygotidae and Slimonidae. Despite their classification as pterygotioids, the hughmilleriids possessed several characteristics shared with other eurypterid groups, such as the lanceolate telson (the most posterior segment of the body). Hughmilleriids are defined as pterygotioid eurypterids with swimming legs similar to those of the type genus, ''Hughmilleria'' (that is, 7th and 8th leg segments narrow and 9th segment very small), and whose second to fifth pair of appendages were spiniferous. Some further diagnostic characters unite the group, such as the slightly enlarged chelicerae (frontal appendages) and the streamlined shape of their b ...
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Pterygotioidea
Pterygotioidea (the name deriving from the type genus ''Pterygotus'', meaning "winged one") is a superfamily of eurypterids, an extinct group of aquatic arthropods. Pterygotioids were the most derived members of the infraorder Diploperculata and the sister group of the adelophthalmoid eurypterids. The group includes the basal and small hughmilleriids, the larger and specialized slimonids and the famous pterygotids which were equipped with robust and powerful cheliceral claws. Though the more primitive hughmilleriids were small, '' Hughmilleria wangi'' being the smallest of all pterygotioids at just in length, later members of the group, particularly in the Pterygotidae, would become the largest known arthropods to ever exist with several genera surpassing in length. Among all currently recognized eurypterid clades, the Pterygotioidea is the most diverse, containing over 50 species in 10 genera. With the number of recognized eurypterid species being around 250, pterygotio ...
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Herefordopterus
''Herefordopterus'' is a genus of eurypterid, an extinct group of aquatic arthropods. ''Herefordopterus'' is classified as part of the family Hughmilleriidae, a basal family in the highly derived Pterygotioidea superfamily of eurypterids. Fossils of the single and type species, ''H. banksii'', have been discovered in deposits of Silurian age in Herefordshire and Shropshire, England. The genus is named after Herefordshire, where most of the ''Herefordopterus'' fossils have been found. The specific epithet honors Richard Banks, who found several well-preserved specimens, including the first ''Herefordopterus'' fossils. ''Herefordopterus'' is classified in Hughmilleriidae, a pterygotioid family that is differentiated by their streamlined bodies, the enlargement of its medium-sized chelicerae and the presence of paired spines on the walking appendages. It was distinguished for combining characteristics of '' Hughmilleria'' with those of the derived pterygotioids, showing a more adva ...
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Hughmilleria
''Hughmilleria'' is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of ''Hughmilleria'' have been discovered in deposits of the Silurian age in China and the United States. Classified as part of the basal family Hughmilleriidae, the genus contains three species, ''H. shawangunk'' from the eastern United States, ''H. socialis'' from Pittsford, New York, and ''H. wangi'' from Hunan, China. The genus is named in honor of the Scottish geologist Hugh Miller. ''H. socialis'' is the type species of Hughmilleriidae, a eurypterid family classified in the superfamily Pterygotioidea that is differentiated by their streamlined bodies, the enlargement of their medium-sized chelicerae and the presence of paired spines on the walking appendages. With the biggest specimen measuring 20 centimetres (8 inches) in length, ''Hughmilleria'' is considered a eurypterid of small size. Description ''Hughmilleria'' is the most basal (primitive) known member of the Pterygotioidea. ...
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Slimonidae
Slimonidae (the name deriving from the type genus '' Slimonia'', which is named in honor of Welsh fossil collector and surgeon Robert Slimon) is a family of eurypterids, an extinct group of aquatic arthropods. Slimonids were members of the superfamily Pterygotioidea and the family most closely related to the derived pterygotid eurypterids, which are famous for their cheliceral claws and great size. Many characteristics of the Slimonidae, such as their flattened and expanded telsons (the posteriormost division of their bodies), support a close relationship between the two groups. Slimonids are defined as pterygotioid eurypterids with swimming legs similar to those of the type genus, ''Slimonia'', and the second to fifth pair of appendages being non-spiniferous. The family contains only two genera, the almost completely known ''Slimonia'' and '' Salteropterus'', which is known only from the telson and the metastoma (a large plate part of the abdomen). Both slimonid genera preserv ...
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Slimonid
Slimonidae (the name deriving from the type genus '' Slimonia'', which is named in honor of Welsh fossil collector and surgeon Robert Slimon) is a family of eurypterids, an extinct group of aquatic arthropods. Slimonids were members of the superfamily Pterygotioidea and the family most closely related to the derived pterygotid eurypterids, which are famous for their cheliceral claws and great size. Many characteristics of the Slimonidae, such as their flattened and expanded telsons (the posteriormost division of their bodies), support a close relationship between the two groups. Slimonids are defined as pterygotioid eurypterids with swimming legs similar to those of the type genus, ''Slimonia'', and the second to fifth pair of appendages being non-spiniferous. The family contains only two genera, the almost completely known ''Slimonia'' and ''Salteropterus'', which is known only from the telson and the metastoma (a large plate part of the abdomen). Both slimonid genera preserve ...
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Pterygotidae
Pterygotidae (the name deriving from the type genus ''Pterygotus'', meaning "winged one") is a family of eurypterids, an extinct group of aquatic arthropods. They were members of the superfamily Pterygotioidea. Pterygotids were the largest known arthropods to have ever lived with some members of the family, such as ''Jaekelopterus'' and '' Acutiramus'', exceeding 2 metres (6.6 ft) in length. Their fossilized remains have been recovered in deposits ranging in age from 428 to 372 million years old ( Late Silurian to Late Devonian). One of the most successful groups of eurypterids, the pterygotids were the only eurypterid family to achieve a truly worldwide distribution. Several evolutionary innovations made the pterygotids unique among the eurypterids, with large and flattened telsons (the posteriormost division of the body) likely used as rudders to provide additional agility and enlarged chelicerae (frontal appendages) with claws. These claws were robust and possessed tee ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs (book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-producing ...
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Chelicerata
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including Opiliones, harvestmen, scorpions, spiders, Solifugae, solifuges, Parasitiformes, ticks, and Acariformes, mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from Burgess Shale, 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of Pycnogonida, pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arth ...
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Adelophthalmoidea
Adelophthalmidae (the name deriving from the type genus ''Adelophthalmus'', meaning "no obvious eyes") is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina. Adelophthalmid eurypterids were small and swimming eurypterids that appeared in the Silurian period. With the earliest known members of the group, '' Nanahughmilleria prominens'' and '' Parahughmilleria maria'', being known from deposits of Early Silurian (possibly the Llandovery epoch) age and the last members, belonging to the long-lasting and widespread genus ''Adelophthalmus'', going extinct in the Cisuralian, Early Permian, the Adelophthalmidae is the longest lasting single family of eurypterids. The survival of the group, and of swimming eurypterids (the suborder Eurypterina) beyond the Late Devonian is entirely due to the ...
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Prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups.) The word ''cephalothorax'' is derived from the Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. Arachnid anatomy Fovea The fovea is the centre of the cephalothorax and is located behind the head (only in spiders).Dalton, Steve (2008). ''Spiders; The Ultimate Predators''. A & C Black, London. P.p. 19. . It is often important in identification. It can be transverse or procurved Smith, A. M. (1990c). Baboon spiders: Tarantulas of Afr ...
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