Hesperogavialis
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Hesperogavialis
''Hesperogavialis'' is an extinct genus of gryposuchine gavialid. Fossils have been found from Venezuela and Brazil that date back to the Middle to Late Miocene. Although ''Hesperogavialis'' is one of the best known gavialoids from South America, the posterior portion of the skull is still unknown, making any attempts at classification within the family somewhat more difficult than other gavialoids in which much of the skull is present. The genus possibly comprises three species. The type species, ''H. cruxenti'', has been found in the Urumaco Formation in Venezuela. A second possible species, named ''H. bocquentini'', has been described from the Solimões Formation in Acre, Brazil, and can be distinguished from ''H. cruxenti'' by the asymmetry seen in the anterior portion of the nasals and the small distance between alveoli. A third species can be recognized from the same locality in Acre, although a formal name has yet to be given to it. Phylogenetics Although considered a ...
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Gryposuchinae
Gryposuchinae is an extinct subfamily (biology), subfamily of Gavialidae, gavialid crocodylians. Gryposuchines lived mainly in the Miocene of South America. However, ''Ikanogavialis, "Ikanogavialis" papuensis'' may have survived more recently, into the Late Pleistocene/Holocene. Most were long-snouted coastal forms. The group was named in 2007 and includes genera such as ''Gryposuchus'' and ''Aktiogavialis'', although a 2018 study indicates that the group might be paraphyletic and rather an evolutionary grade towards the gharial. Description Gryposuchines have long, narrow snouts and protruding eye sockets. One distinguishing feature of the group is the lack of a large exposure of the prootic bone around the trigeminal foramen, a hole in the side of the braincase wall. Classification Gryposuchinae was named in 2007 as a subfamily of closely related gavialid crocodilians. It was cladistics, cladistically defined as a stem-based taxon including ''Gryposuchus jessei'' and all croco ...
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Huayquerian
The Huayquerian ( es, Huayqueriense) age is a period of geologic time (9.0–6.8 Ma) within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification. It follows the Mayoan and precedes the Montehermosan age. Etymology The age is named after the Huayquerías Formation in the western Cuyo Basin of northwestern Argentina that was later dated to the Montehermosan. The most complete Huayquerian fauna is found in the Cerro Azul Formation The Cerro Azul Formation ( es, Formación Cerro Azul), in the Buenos Aires Province also described as Epecuén Formation, is a geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in the Colorado Basin ..., in Buenos Aires Province also referred to as Epecuén Formation. Formations Fossil content Correlations Notes and references Notes References Bibliography ;Huayquerías Formation * * ;Andalhuala Formation * ;Camacho Formation * * ...
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Chasicoan
The Chasicoan ( es, Chasiquense or es, Chasicoense) age is a period of geologic time from 10–9 Ma within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Mayoan and precedes the Huayquerian age.Chasicoan
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Etymology

The Chasicoan is named after the
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Urumaco Formation
The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location The Urumaco formation is located in the Urumaco region in the Caribbean coastal Falcón state. The deposits date from 10 to 5.3 million years ago and the Urumaco formation was deposited in an area with large rivers, swamps, estuaries, lagoons and shallow coastal seas. These conditions in the Late Miocene contrast strongly with the current dry environment in the area today. Fauna Cartilaginous fish There are 21 known species of cartilaginous fishes from the Urumaco Formation, belonging to the orders Lamniformes, Carcharhiniformes, Myliobatiformes and Rajiformes. ''Carcharhinus caquetius'' is an endemic species of predator shark from Urumaco. A large number of well-preserved fossils of the sawfish ''Pristis rostra'' have been found in th ...
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Gavialidae
Gavialidae is a family (biology), family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two extant taxon, living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a Generalist and specialist species, generalist diet with mature adults preying upon larger vertebrates, such as Ungulate, ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams (zoologist), Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Nasal Bone
The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Each has two surfaces and four borders. Structure The two nasal bones are joined at the midline internasal suture and make up the bridge of the nose. Surfaces The ''outer surface'' is concavo-convex from above downward, convex from side to side; it is covered by the procerus and nasalis muscles, and perforated about its center by a foramen, for the transmission of a small vein. The ''inner surface'' is concave from side to side, and is traversed from above downward, by a groove for the passage of a branch of the nasociliary nerve. Articulations The nasal articulates with four bones: two of the cranium, the frontal and ethmoid, and two of the face, the opposite nasal and the maxilla. Other animals In primitive bony fish and tetrapod ...
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Neogene Venezuela
The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans (''Homo habilis'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific to the ...
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Fossils Of Brazil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolut ...
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Neogene Brazil
The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans ('' Homo habilis'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific ...
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Prehistoric Pseudosuchian Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Miocene Reptiles Of South America
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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