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Herbst Corpuscle
The corpuscles of Herbst or Herbst corpuscles are nerve-endings similar to the Pacinian corpuscle, found in the mucous membrane of the tongue, in pits on the beak and in other parts of the bodies of birds. They differ from Pacinian corpuscles in being smaller and more elongated, in having thinner and more closely placed capsules, and in that the axis-cylinder in the central clear space is encircled by a continuous row of nuclei. They are named after the German embryologist Curt Alfred Herbst. In many wading birds, a large number of Herbst corpuscles are found embedded in pits on the mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ... that are believed to enable birds to sense prey under wet sand or soil. References External links Description at sciencedaily.com* ...
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Pacinian Corpuscle
Pacinian corpuscle or lamellar corpuscle or Vater-Pacini corpuscle; is one of the four major types of mechanoreceptors (specialized nerve ending with adventitious tissue for mechanical sensation) found in mammalian skin. This type of mechanoreceptor is found in both glabrous (hairless) and hirsute (hairy) skins, viscera, joints and attached to periosteum of bone, primarily responsible for sensitivity to vibration. Few of them are also sensitive to quasi-static or low frequency pressure stimulus. Most of them respond only to sudden disturbances and are especially sensitive to vibration of few hundreds of Hz. The vibrational role may be used for detecting surface texture, e.g., rough vs. smooth. Most of the Pacinian corpuscles act as rapidly adapting mechanoreceptors. Groups of corpuscles respond to pressure changes, e.g. on grasping or releasing an object. Structure Pacinian corpuscles are larger and fewer in number than Meissner's corpuscle, Merkel cells and Ruffini's corpuscl ...
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Mucous Membrane
A mucous membrane or mucosa is a membrane that lines various cavities in the body of an organism and covers the surface of internal organs. It consists of one or more layers of epithelial cells overlying a layer of loose connective tissue. It is mostly of endodermal origin and is continuous with the skin at body openings such as the eyes, eyelids, ears, inside the nose, inside the mouth, lips, the genital areas, the urethral opening and the anus. Some mucous membranes secrete mucus, a thick protective fluid. The function of the membrane is to stop pathogens and dirt from entering the body and to prevent bodily tissues from becoming dehydrated. Structure The mucosa is composed of one or more layers of epithelial cells that secrete mucus, and an underlying lamina propria of loose connective tissue. The type of cells and type of mucus secreted vary from organ to organ and each can differ along a given tract. Mucous membranes line the digestive, respiratory and reproductive trac ...
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Tongue
The tongue is a muscular organ (anatomy), organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive system, digestive process, and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste buds housed in numerous lingual papillae. It is sensitive and kept moist by saliva and is richly supplied with nerves and blood vessels. The tongue also serves as a natural means of oral hygiene, cleaning the teeth. A major function of the tongue is the enabling of speech in humans and animal communication, vocalization in other animals. The human tongue is divided into two parts, an oral cavity, oral part at the front and a pharynx, pharyngeal part at the back. The left and right sides are also separated along most of its length by a vertical section of connective tissue, fibrous tissue (the lingual septum) that results in a groove, the median sulcus, on the tongue's surface. There are two groups of muscle ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the bee hummingbird to the ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming. B ...
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Cell Nucleus
The cell nucleus (pl. nuclei; from Latin or , meaning ''kernel'' or ''seed'') is a membrane-bound organelle found in eukaryotic cells. Eukaryotic cells usually have a single nucleus, but a few cell types, such as mammalian red blood cells, have no nuclei, and a few others including osteoclasts have many. The main structures making up the nucleus are the nuclear envelope, a double membrane that encloses the entire organelle and isolates its contents from the cellular cytoplasm; and the nuclear matrix, a network within the nucleus that adds mechanical support. The cell nucleus contains nearly all of the cell's genome. Nuclear DNA is often organized into multiple chromosomes – long stands of DNA dotted with various proteins, such as histones, that protect and organize the DNA. The genes within these chromosomes are structured in such a way to promote cell function. The nucleus maintains the integrity of genes and controls the activities of the cell by regulating gene expres ...
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Curt Alfred Herbst
Curt Alfred Herbst (29 May 1866 – 9 May 1946) was a German embryologist and zoologist who studied development of cells and tissues. He noted the mechanosensory organs now known as '' Herbst corpuscles'' in the bills of birds. Herbst was born to Heinrich and Henriette Martin in Meuselwitz, Thuringia. He was educated in Geneva and Jena where his teachers included Carl Vogt and Ernst Haeckel. He also went to the Polytechnic in Zurich to study chemistry. He studied echinoderm larvae and noted the effect of lithium ion gradients on morphogenesis and noted sex-determination in ''Bonellia viridis'' through environmental concentrations of ions. After obtaining a Ph.D. in 1889 he visited Southeast Asia along with Hans Driesch, visiting Ceylon, Java and the Trieste marine station. He then went to Heidelberg to study under Otto Bütschli and became an associate professor in 1906. He received an honorary doctorate from the University of Halle. His students included Helmuth Plessner Helmu ...
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Charadriformes
Charadriiformes (, from ''Charadrius'', the type genus of family Charadriidae) is a diverse order of small to medium-large birds. It includes about 390 species and has members in all parts of the world. Most charadriiform birds live near water and eat invertebrates or other small animals; however, some are pelagic (seabirds), others frequent deserts, and a few are found in dense forest. Members of this group can also collectively be referred to as shorebirds. Taxonomy, systematics and evolution The order was formerly divided into three suborders: * The waders (or "Charadrii"): typical shorebirds, most of which feed by probing in the mud or picking items off the surface in both coastal and freshwater environments. * The gulls and their allies (or " Lari"): these are generally larger species which take fish from the sea. Several gulls and skuas will also take food items from beaches, or rob smaller species, and some have become adapted to inland environments. * The auks (or "Alcae ...
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Mandible
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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