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Helicoprionidae
The Helicoprionidae, also known as the Agassizodontidae are an extinct, poorly known family of bizarre holocephalids within the poorly understood order Eugeneodontida. Members of the Helicoprionidae possessed a unique "tooth-whorl" on the symphysis of the lower jaw and pectoral fins supported by long radials. The closest living relatives of the Helicoprionidae and all other eugeneodontids are the ratfishes. The anatomy of the tooth-whorl differed amongst genus and species, some possessing complete spirals (such as those of ''Helicoprion ''Helicoprion'' is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extin ...''), others possessing halved spirals (seen in '' Parahelicoprion''), and some with wedged half-spirals (seen in '' Sarcoprion''). Each tooth-whorl is thought to be adapted to a different type of p ...
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Agassizodontidae
The Helicoprionidae, also known as the Agassizodontidae are an extinct, poorly known family of bizarre holocephalids within the poorly understood order Eugeneodontida. Members of the Helicoprionidae possessed a unique "tooth-whorl" on the symphysis of the lower jaw and pectoral fins supported by long radials. The closest living relatives of the Helicoprionidae and all other eugeneodontids are the ratfishes. The anatomy of the tooth-whorl differed amongst genus and species, some possessing complete spirals (such as those of ''Helicoprion''), others possessing halved spirals (seen in ''Parahelicoprion ''Parahelicoprion'' is an extinct genus of shark-like eugeneodontid holocephalids from the Permian of the Ural Mountains (Russia) and Copacabana Formation, Bolivia. The genus name, from "nearly coiled saw" in Greek, directly refers to ''Helico ...''), and some with wedged half-spirals (seen in '' Sarcoprion''). Each tooth-whorl is thought to be adapted to a different type of pre ...
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Helicoprion
''Helicoprion'' is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of ''Helicoprion'' are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian (Early Permian) to the Roadian stage of the Guadalupian (Middle Permian). The closest living relatives of ''Helicoprion'' (and other eugeneodonts) are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of ''Helicoprion'' via morphometric analysis of the tooth whorls found only ''H. davisii, H. bessonowi'' ...
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Eugeneodontida
The Eugeneodontida is an extinct and poorly known order of cartilaginous fishes. They possessed "tooth-whorls" on the symphysis of either the lower or both jaws and pectoral fins supported by long radials. They probably lacked pelvic fins and anal fins. The palatoquadrate was either fused to the skull or reduced. Now determined to be within the Holocephali, their closest living relatives are ratfish. The eugeneodonts are named after paleontologist Eugene S. Richardson, Jr. The Eugeneodontida disappeared in the Early Triassic. Members of the Eugeneodontida are further classified into different families, the most well-preserved members that have been discovered are commonly placed within the families Helicoprionidae ("spiral saws"), and Edestidae ("those which devour"), the former containing the genera ''Helicoprion'', '' Sarcoprion'', and '' Parahelicoprion'', and the latter containing the genera '' Edestus'', ''Lestrodus'', and '' Metaxyacanthus''. All eugeneodonts are thought ...
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Agassizodus
''Agassizodus'' is an extinct genus of eugeneodont holocephalan from the Carboniferous. It belongs to the family Helicoprionidae, which is sometimes called Agassizodontidae. Like other members of its family, it possessed a symphyseal tooth whorl, which was likely present at the tip of the lower jaw and associated with lateral crushing toothplates. The type species, ''A. variabilis'', was originally named ''Lophodus variabilis'' until the name "Lophodus" was determined to be preoccupied. ''A. variabilis'' was originally based on tooth fragments from the Pennsylvanian of Illinois, but the authors who named the genus (St. John and Worthen, 1875) also referred a massive jaw from Osage County, Kansas Osage County (county code OS) is a county located in the U.S. state of Kansas. As of the 2020 census, the county population was 15,766. Its county seat is Lyndon, and its most populous city is Osage City. The county was originally organized i .... The Osage jaw shared some simi ...
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Sarcoprion
''Sarcoprion'' (from the Ancient Greek for "flesh saw") is an extinct genus of eugeneodontid holocephalids from the Permian of Greenland. Similar to other eugeneodontids such as ''Edestus'' and ''Helicoprion'', it was best known for its extremely bizarre tooth morphology compared to other species of sharks and their closest relatives, the chimaeras. Compared to other members of the Helicoprionidae (teeth of Agassiz), its "tooth whorls" were found to be sharper, more compact, and in better condition than other sharks of the time, and refrained from growing to extremely unwieldy forms that would raise questions about its ability to feed properly. The genus contains one species, ''Sarcoprion edax'' ("gluttonous flesh saw"), found in Permian-aged marine strata of ''Meddelelser om Grønland''. Description ''Sarcoprion'' had a jaw and mouth structure which allowed it to be more hydrodynamic, reducing the size and shape of the tooth whorl and increasing the size of the rostrum. ''Sarc ...
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Early Carboniferous
Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early Branch, a stream in Missouri * Early County, Georgia Other uses * Early (Scritti Politti album), ''Early'' (Scritti Politti album), 2005 * Early (A Certain Ratio album), ''Early'' (A Certain Ratio album), 2002 * Early (name) * Early effect, an effect in transistor physics * Early Records, a record label * the early part of the morning See also

* Earley (other) {{disambiguation, geo ...
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Toxoprion
''Toxoprion'' (Ancient Greek for "bow saw") is an extinct genus of eugeneodont holocephalids whose fossils are found in marine strata from the Early Carboniferous until the Late Permian near Eureka, Nevada. Description ''Toxoprion'' was one of the many Carboniferous eugeneodonts which bore a palatoquadrate fused to its skull or reduced in other forms, and had its heavily serrated teeth grow outwards on the symphysis of the lower jaw similar to a rounded saw. Despite its jaw showing similarities to another eugeneodont, ''Helicoprion'', the tooth row of ''Toxoprion'' does not coil back on itself, as seen in ''Helicoprion'' but instead makes a downward curved tooth whorl similar to '' Campyloprion'', ''Lestrodus'', and ''Edestus''. As in most members of eugeneodontida, the smaller teeth found near the anterior end of the whorl represent the oldest teeth, which the animal grew as a juvenile and the larger and younger teeth found near the back of the whorl represent teeth grown in ...
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Ratfish
Chimaeras are cartilaginous fish in the order Chimaeriformes , known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively. At one time a "diverse and abundant" group (based on the fossil record), their closest living relatives are sharks and rays, though their last common ancestor with them lived nearly 400 million years ago. Today, they are largely confined to deep water. Description and habits Chimaeras live in temperate ocean floors down to deep, with few occurring at depths shallower than . Exceptions include the members of the genus ''Callorhinchus'', the rabbit fish and the spotted ratfish, which locally or periodically can be found at shallower depths. Consequently, these are also among the few species from the chimaera order kept in public aquaria. They live in all the oceans except for the Arctic and Antarctic oceans. They have elongated, soft bod ...
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Symphysis
A symphysis (, pl. symphyses) is a fibrocartilaginous fusion between two bones. It is a type of cartilaginous joint, specifically a secondary cartilaginous joint. # A symphysis is an amphiarthrosis, a slightly movable joint. # A growing together of parts or structures. Unlike synchondroses, symphyses are permanent. Examples The more prominent symphyses are: * the pubic symphysis * sacrococcygeal symphysis * intervertebral disc between two vertebrae * in the sternum, between the manubrium and body Body may refer to: In science * Physical body, an object in physics that represents a large amount, has mass or takes up space * Body (biology), the physical material of an organism * Body plan, the physical features shared by a group of anima ... * mandibular symphysis, in the jaw Symphysis disorders Pubic symphysis diastasis Pubic symphysis diastasis, is an extremely rare complication that occurs in women who are giving birth. Separation of the two pubic bones during deli ...
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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Holocephali
Holocephali ("complete heads"), sometimes given the term Euchondrocephali, is a subclass of cartilaginous fish in the class Chondrichthyes. The earliest fossils are of teeth and come from the Devonian period. Little is known about these primitive forms, and the only surviving group in the subclass is the order Chimaeriformes. Chimaeriformes, commonly known as chimaeras, includes the rat fishes in the genus '' Chimaera'', and the elephant fishes in the genus ''Callorhinchus''. These fishes move by using sweeping movements of their large pectoral fins. They are deep sea fish with slender tails, living close to the seabed to feed on benthic invertebrates. They lack a stomach, their food moving directly into the intestine. Extinct holocephalans were much more diverse in lifestyles, including shark-like predatory forms and slow, durophagous fish. Characteristics Members of this taxon preserve today some features of elasmobranch life in Paleozoic times, though in other respects th ...
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