Graphis Tetralocularis
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Graphis Tetralocularis
''Graphis tetralocularis'' is a species of crustose lichen in the family Graphidaceae. Found in Rwanda, it was formally described as a new species in 2005 by Christina Bock and Markus Hauck. The type specimen was collected by the first author from Akagera National Park at an altitude of ; here it was found growing on a twig of a ''Nuxia floribunda'' tree. It is only known to occur at the type locality, which is a dry forest dominated by the trees ''Nuxia floribunda'', ''Haplocoelum gallense'', and ''Strychnos usambarensis''. The lichen has a thin, smooth, whitish grey to grey-green crustose thallus. The specific epithet ''tetralocularis'' refers to the unusual morphology of its four-chambered ascospore An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or s ...s. ''Graphis tetralocularis' ...
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Crustose Lichen
Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cortex (botany), cortex layer, an algal layer, and a medulla. The upper cortex layer is differentiated and is usually pigmented. The algal layer lies beneath the cortex. The medulla fastens the lichen to the substrate and is made up of Fungus, fungal hyphae. The surface of crustose lichens is characterized by branching cracks that periodically close in response to climatic variations such as alternate wetting and drying regimes. Subtypes * Powdery – considered as the simplest subtype due to the absence of an organized thallus. :The thallus appears powdery. :E.g. Genera ''Lepraria'', ''Vezdaea'' * Endolithic – grows inside the rock, usually in interstitial spaces between mineral grains. The :upper cortex is usually d ...
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Botanical Name
A botanical name is a formal scientific name conforming to the '' International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the ''International Code of Nomenclature for Cultivated Plants'' (ICNCP). The code of nomenclature covers "all organisms traditionally treated as algae, fungi, or plants, whether fossil or non-fossil, including blue-green algae ( Cyanobacteria), chytrids, oomycetes, slime moulds and photosynthetic protists with their taxonomically related non-photosynthetic groups (but excluding Microsporidia)." The purpose of a formal name is to have a single name that is accepted and used worldwide for a particular plant or plant group. For example, the botanical name ''Bellis perennis'' denotes a plant species which is native to most of the countries of Europe and the Middle East, where it has accumulated various names in many languages. Later, the plant was intro ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Graphis (lichen)
''Graphis'' is a genus of lichenized fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ... in the family Graphidaceae. Selected species *'' Graphis alboscripta'' *'' Graphis albotecta'' *'' Graphis alpestris'' *'' Graphis analoga'' *'' Graphis anfractuosa'' *'' Graphis antillarum'' *'' Graphis aperiens'' *'' Graphis apertella'' *'' Graphis appendiculata'' *'' Graphis archeri'' *'' Graphis breussii'' *'' Graphis bulacana'' *'' Graphis caesiella'' *'' Graphis carassensis'' *'' Graphis caribica'' *'' Graphis centrifuga'' *'' Graphis cervina'' *'' Graphis chlorotica'' *'' Graphis chrysocarpa'' *'' Graphis cincta'' *'' Graphis cinerea'' *'' Graphis componens'' *'' Graphis conturbata'' *'' Graphis crassilabra'' *'' Graphis crebra'' *'' Graphis crystallifera'' *'' Graphis cycasicol ...
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Thin-layer Chromatography
Thin-layer chromatography (TLC) is a chromatography technique used to separate non-volatile mixtures. Thin-layer chromatography is performed on a sheet of an inert substrate such as glass, plastic, or aluminium foil, which is coated with a thin layer of adsorbent material, usually silica gel, aluminium oxide (alumina), or cellulose. This layer of adsorbent is known as the stationary phase. After the sample has been applied on the plate, a solvent or solvent mixture (known as the mobile phase) is drawn up the plate via capillary action. Because different analytes ascend the TLC plate at different rates, separation is achieved. It may be performed on the analytical scale as a means of monitoring the progress of a reaction, or on the preparative scale to purify small amounts of a compound. TLC is an analytical tool widely used because of its simplicity, relative low cost, high sensitivity, and speed of separation. TLC functions on the same principle as all chromatography: a com ...
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Secondary Metabolite
Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the normal growth, development, or reproduction of the organism. Instead, they generally mediate ecological interactions, which may produce a selective advantage for the organism by increasing its survivability or fecundity. Specific secondary metabolites are often restricted to a narrow set of species within a phylogenetic group. Secondary metabolites often play an important role in plant defense against herbivory and other interspecies defenses. Humans use secondary metabolites as medicines, flavourings, pigments, and recreational drugs. The term secondary metabolite was first coined by Albrecht Kossel, a 1910 Nobel Prize laureate for medicine and physiology in 1910. 30 years later a Polish botanist Friedrich Czapek described secondary metabolit ...
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Atranorin
Atranorin is a chemical substance produced by some species of lichen. It is a secondary metabolite belonging to a group of compounds known as depsides. Atranorin has analgesic, anti-inflammatory, antibacterial, antifungal, cytotoxic, antioxidant, antiviral, and immunomodulatory properties. In rare cases, people can react allergic Allergies, also known as allergic diseases, refer a number of conditions caused by the hypersensitivity of the immune system to typically harmless substances in the environment. These diseases include hay fever, food allergies, atopic derma ... to atranorin. References Further reading * * * * Polyphenols Lichen products {{organic-compound-stub ...
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Ascospore
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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Graphidaceae
The Graphidaceae are a family of lichens in the order Ostropales. Distribution and ecology The vast majority of Graphidaceae species are restricted to the tropics. Most Graphidaceae species are epiphytic (i.e. they grow only on plants). Genera A recent (2020) estimates places 31 genera and about 990 species in Graphidaceae. The following list indicates the genus name, the taxonomic authority, year of publication, and the number of species: *'' Acanthothecis'' – 5 spp. *'' Acanthotrema'' – 1 sp. *'' Aggregatorygma'' – 1 sp. *'' Allographa'' – 183 spp. *'' Amazonotrema'' – 1 sp. *'' Ampliotrema'' – 1 sp. *'' Anomalographis'' – 2 spp. *'' Anomomorpha'' – 8 spp. *'' Astrochapsa'' – 29 spp. *'' Austrotrema'' – 3 spp. *'' Borinquenotrema'' – 1 sp. *'' Byssotrema'' – 1 sp. *'' Carbacanthographis'' – 22 spp. *'' Chapsa'' – 51 spp. *'' Chroodiscus'' – 17 spp. *'' Clandestinotrema'' – 17 spp. *'' Compositrema'' – 4 spp. *'' Corticorygma' ...
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Strychnos Usambarensis
''Strychnos usambarensis'' is a shrub or small tree up to 15m tall or a 70m long liane of Sub-Saharan Africa, occurring in forest and woodland, mountain ravines and coastal bush, often on rocky slopes and named for the Usambara Mountains of Tanzania. The species is found from Guinea east to Nigeria, from Congo east to Kenya and south to kloofs of the Magaliesberg of South Africa. Bantu tribes from Rwanda and Tanzania produce an arrow poison from the root bark and leaves of this species, sometimes combining it with extracts from other plants. (see Toxalbumin) The leaves are opposite, held in a horizontal plane, ovate to elliptic in shape with a characteristic drip-tip, and glossy dark green above. Flowers are small, in axillary clusters, whitish to yellowish-green, and fragrant. Fruits are soft-shelled and small, 1-1.5 cm in diameter, yellow when ripe, tapering to a collar at peduncle. The genus ''Strychnos'' is represented by some 300 species of lianes, shrubs and small tre ...
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Haplocoelum Gallense
''Haplocoelum'' is a genus of plants in the family Sapindaceae The Sapindaceae are a family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1858 accepted species. Examples include horse chestnut, maples, ackee and lychee. The Sapindaceae occur in tempera .... It contains the following species (but this list may be incomplete): Sapindaceae genera Taxonomy articles created by Polbot {{sapindales-stub ...
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