Gollum (genus)
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Gollum (genus)
''Gollum'' is a genus of ground sharks in the family Pseudotriakidae, native to the southwestern Pacific Ocean. The genus was described in 1973 by biologist Leonard Compagno, who named it named after the character Gollum from J. R. R. Tolkien's works, noting the species ''Gollum attenuatus'' (the slender smooth-hound) "bears some resemblance in form and habits". Species The currently described species and one undescribed species in this genus are: * ''G. attenuatus'' (Garrick, 1954) (slender smooth-hound) * ''G. suluensis'' Last & Gaudiano, 2011 ( Sulu gollumshark) * ''Gollum'' sp. B not yet described (white-marked gollumshark) See also * ''Aenigmachanna gollum ''Aenigmachanna gollum'', the Gollum snakehead, is a species of aquifer-dwelling dragon snakehead fish that is endemic to the Indian state of Kerala. Description ''A. gollum'' has an elongated, eel-like body coloured mostly brown and beige. ...'', a species of fish named after the same character from Tolkien's ...
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Slender Smooth-hound
The slender smooth-hound or gollumshark (''Gollum attenuatus'') is a species of ground shark in the family Pseudotriakidae. It is endemic to the waters around New Zealand, where it is usually found close to the bottom over the continental slope at depths of . An extremely slim, plain brownish shark reaching in length, the slender smooth-hound can be identified by its broad, flattened head with a long, distinctively bell-shaped snout. Its mouth is angular with short furrows at the corners, and contains a very high number of tooth rows in both jaws. Its two dorsal fins are roughly equal in size. The diet of the slender smooth-hound is diverse, but dominated by small, benthic bony fishes and decapod crustaceans. It exhibits a specialized form of aplacental viviparity with oophagy: the females produce a single capsule in each uterus that contains 30–80 ova, of which one ovum develops into an embryo that consumes the rest of the ova and stores the yolk material in its external y ...
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Leonard Joseph Victor Compagno
Leonard Joseph Victor Compagno is an international authority on shark taxonomy and the author of many scientific papers and books on the subject, best known of which is his 1984 catalogue of shark species produced for the Food and Agriculture Organization (FAO) of the United Nations. Compagno was mentioned in the credits of the 1975 film ''Jaws'' along with the National Geographic Society. Career *Ph.D, Stanford University, 1979 *Adjunct professor, San Francisco State University, 1979 to 1985 *Curator of Fishes in the Division of Life Sciences and Head of the Shark Research Centre (SRC), Iziko Museums, Cape Town *Director, Shark Research Institute(SRI) Selected bibliography *Compagno, L.J.V., 1979. ''Carcharhinoid sharks: morphology, systematics and phylogeny''. Unpublished Ph. D. Thesis, Stanford University, 932 p. Available from University Microfilms International, Ann Arbor, Michigan. *Leonard Compagno, 1984a. FAO The Food and Agriculture Organization of the United Nati ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Carcharhiniformes
Carcharhiniformes , the ground sharks, are the largest order of sharks, with over 270 species. They include a number of common types, such as catsharks, swellsharks, and the sandbar shark. Members of this order are characterized by the presence of a nictitating membrane over the eye, two dorsal fins, an anal fin, and five gill slits. The families in the order Carcharhiniformes are expected to be revised; recent DNA studies show that some of the conventional groups are not monophyletic. The oldest members of the order appeared during the Middle-Late Jurassic, which have teeth and bodyforms that are morphologically similar to living catsharks. Carchariniformes first underwent major diversification during the Late Cretaceous, initially as small-sized forms, before radiating into medium and large body sizes during the Cenozoic. Families According to FishBase, the nine families of ground sharks are:
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Pseudotriakidae
The Pseudotriakidae are a small family of ground sharks, belonging to the order Carcharhiniformes, containing the false catsharks (genera ''Pseudotriakis'' and ''Planonasus'') and gollumsharks (genus ''Gollum''). It contains the only ground shark species that exhibit intrauterine oophagy, in which developing fetus A fetus or foetus (; plural fetuses, feti, foetuses, or foeti) is the unborn offspring that develops from an animal embryo. Following embryonic development the fetal stage of development takes place. In human prenatal development, fetal deve ...es are nourished by eggs produced by their mother. Undescribed species One undescribed species is known - one in the genus ''Gollum'', (''Gollum'' sp. B) - the white-marked gollumshark.Weigmann, S., Stehmann, M.F.W. & Thiel, R. (2013): ''Planonasus parini'' n. g. and n. sp., a new genus and species of false cat sharks (Carchariniformes, Pseudotriakidae) from the deep northwestern Indian Ocean off Socotra Islands. ''Zoot ...
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Pacific Ocean
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continents of Asia and Oceania in the west and the Americas in the east. At in area (as defined with a southern Antarctic border), this largest division of the World Ocean—and, in turn, the hydrosphere—covers about 46% of Earth's water surface and about 32% of its total surface area, larger than Earth's entire land area combined .Pacific Ocean
. '' Britannica Concise.'' 2008: Encyclopædia Britannica, Inc.
The centers of both the

Leonard Compagno
Leonard Joseph Victor Compagno is an international authority on shark taxonomy and the author of many scientific papers and books on the subject, best known of which is his 1984 catalogue of shark species produced for the Food and Agriculture Organization (FAO) of the United Nations. Compagno was mentioned in the credits of the 1975 film ''Jaws'' along with the National Geographic Society. Career *Ph.D, Stanford University, 1979 *Adjunct professor, San Francisco State University, 1979 to 1985 *Curator of Fishes in the Division of Life Sciences and Head of the Shark Research Centre (SRC), Iziko Museums, Cape Town *Director, Shark Research Institute(SRI) Selected bibliography *Compagno, L.J.V., 1979. ''Carcharhinoid sharks: morphology, systematics and phylogeny''. Unpublished Ph. D. Thesis, Stanford University, 932 p. Available from University Microfilms International, Ann Arbor, Michigan. *Leonard Compagno, 1984a. FAO The Food and Agriculture Organization of the United Nati ...
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Gollum
Gollum is a fictional Tolkien's monsters, character in J. R. R. Tolkien's Middle-earth legendarium. He was introduced in the 1937 Fantasy (genre), fantasy novel ''The Hobbit'', and became important in its sequel, ''The Lord of the Rings''. Gollum was a Hobbit#Types, Stoor Hobbit of the River-folk who lived near the Gladden Fields. In ''The Lord of the Rings'' it is stated that he was originally known as Sméagol, corrupted by the One Ring, and later named Gollum after his habit of making "a horrible swallowing noise in his throat". Sméagol obtained the Ring by murdering his relative Déagol, who found it in the River Anduin. Gollum referred to the Ring as "my precious" or "precious", and it extended his life far beyond natural limits. Centuries of the Ring's influence twisted Gollum's body and mind, and, by the time of the novels, he "loved and hated [the Ring], as he loved and hated himself." Throughout the story, Gollum was torn between his lust for the Ring and his desire t ...
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Jack Garrick
John Andrew Frank "Jack" Garrick (1928 – August 30, 2018) was a New Zealand ichthyologist. He specialized in elasmobranchs and published many books and articles about shark and ray biology. In 1982, he published a thorough taxonomy on sharks of the genus ''Carcharhinus'', where he identified the smoothtooth blacktip shark as a new species. He is the species authority for several types of sharks, including the New Zealand lanternshark. Garrick was a zoology professor at Victoria University of Wellington, appointed to a personal chair in 1971. He had a primary interest in the taxonomy of sharks and rays, and carried out the first exploratory deep-sea sampling using specially adapted cone nets, baited traps, and longlines, regularly to depths greater than 2000 m. Many new and rare species were obtained by use of these innovative techniques. He was responsible for the notable discovery of the first New Zealand specimens of orange roughy in 1957 (which subsequently formed the basis ...
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Slender Smooth-hound
The slender smooth-hound or gollumshark (''Gollum attenuatus'') is a species of ground shark in the family Pseudotriakidae. It is endemic to the waters around New Zealand, where it is usually found close to the bottom over the continental slope at depths of . An extremely slim, plain brownish shark reaching in length, the slender smooth-hound can be identified by its broad, flattened head with a long, distinctively bell-shaped snout. Its mouth is angular with short furrows at the corners, and contains a very high number of tooth rows in both jaws. Its two dorsal fins are roughly equal in size. The diet of the slender smooth-hound is diverse, but dominated by small, benthic bony fishes and decapod crustaceans. It exhibits a specialized form of aplacental viviparity with oophagy: the females produce a single capsule in each uterus that contains 30–80 ova, of which one ovum develops into an embryo that consumes the rest of the ova and stores the yolk material in its external y ...
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Peter R
Peter may refer to: People * List of people named Peter, a list of people and fictional characters with the given name * Peter (given name) ** Saint Peter (died 60s), apostle of Jesus, leader of the early Christian Church * Peter (surname), a surname (including a list of people with the name) Culture * Peter (actor) (born 1952), stage name Shinnosuke Ikehata, Japanese dancer and actor * ''Peter'' (album), a 1993 EP by Canadian band Eric's Trip * ''Peter'' (1934 film), a 1934 film directed by Henry Koster * ''Peter'' (2021 film), Marathi language film * "Peter" (''Fringe'' episode), an episode of the television series ''Fringe'' * ''Peter'' (novel), a 1908 book by Francis Hopkinson Smith * "Peter" (short story), an 1892 short story by Willa Cather Animals * Peter, the Lord's cat, cat at Lord's Cricket Ground in London * Peter (chief mouser), Chief Mouser between 1929 and 1946 * Peter II (cat), Chief Mouser between 1946 and 1947 * Peter III (cat), Chief Mouser between 1947 ...
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Sulu Gollumshark
The Sulu gollumshark (''Gollum suluensis'') is a species of ground shark in the family Pseudotriakidae, found off Palawan Island in the southern Philippines. This recently discovered species was discovered at a fish market in Palawan during a project led by the World Wildlife Fund during the 1990s to investigate elasmobranch biodiversity in the area. The Sulu gollumshark derives its name from the Sulu Sea in the Philippines. They can grow to sizes of 58.5 cm (1.9 ft) in length, and feature darker coloration and a wider head relative to the closely related slender smooth-hound The slender smooth-hound or gollumshark (''Gollum attenuatus'') is a species of ground shark in the family Pseudotriakidae. It is endemic to the waters around New Zealand, where it is usually found close to the bottom over the continental slope ..., another species of gollumshark. References Gollum (genus) Taxa named by Peter R. Last Fish described in 2011 {{shark-stub ...
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