Graphis Marusae
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Graphis Marusae
''Graphis marusae'' is a species of corticolous lichen, corticolous (bark-dwelling) crustose lichen in the family Graphidaceae. It is found in a relict tropical lowland rainforest in Veracruz, Mexico, growing in exposed understory. Taxonomy The lichen-forming fungus was species description, described as new to science in 2011 by the lichenologists Alejandrina Peña and Robert Lücking. The type (biology), type specimen was collected by Bárcenas Peña from the Los Tuxtlas Biosphere Reserve at an elevation of , where it was growing in a lowland rainforest on the bark of ''Astrocaryum mexicanum'' and ''Pseudolmedia oxyphyllaria''. It is characterised by its green thallus, its 1–5 mm-long (elongated and slit-like fruiting bodies) with grey-black labia. Description The thallus of ''Graphis marusae'' reaches up to in diameter with a thickness of 50–100 μm. The surface of the thallus is smooth, shiny, and pale greenish-grey in colour. There is no prothallus#In lichens, prot ...
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Robert Lücking
Robert Lücking (born 1964) is a German lichenologist. He is a leading expert on foliicolous lichens–lichens that live on leaves. Life and career Born in Ulm in 1964, Lücking earned both his master's (1990) and PhD degree (1994) at the University of Ulm. Both degrees concerned the taxonomy, ecology, and biodiversity of foliicolous lichens. His graduate supervisor was mycologist and bryologist Sieghard Winkler, who had previously studied epiphyllous (upper leaf-dwelling) fungi in El Salvador and Colombia. In 1996 Lücking was awarded the Mason E. Hale award for an "outstanding doctoral thesis presented by a candidate on a lichenological theme". His thesis was titled ''Foliikole Flechten und ihre Mikrohabitatpraferenzen in einem tropischen Regenwald in Costa Rica'' ("Foliicolous lichens and their microhabitat preferences in a tropical rainforest in Costa Rica"). In this work, Lücking recorded 177 foliicolous lichen species from the shrub layer in a Costa Rican tropical forest. L ...
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Prothallus
A prothallus, or prothallium, (from Latin ''pro'' = forwards and Greek ''θαλλος'' (''thallos'') = twig) is usually the gametophyte stage in the life of a fern or other pteridophyte. Occasionally the term is also used to describe the young gametophyte of a liverwort or peat moss as well. In lichens it refers to the region of the thallus that is free of algae. The prothallus develops from a germinating spore. It is a short-lived and inconspicuous heart-shaped structure typically 2–5 millimeters wide, with a number of rhizoids (root-like hairs) growing underneath, and the sex organs: archegonium (female) and antheridium (male). Appearance varies quite a lot between species. Some are green and conduct photosynthesis while others are colorless and nourish themselves underground as saprotrophs. Alternation of generations Spore-bearing plants, like all plants, go through a life-cycle of alternation of generations. The fully grown sporophyte, what is commonly referred to as ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichens Described In 2011
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Graphis (lichen)
''Graphis'' is a genus of lichenized fungi in the family Graphidaceae. Selected species *''Graphis alboscripta'' *''Graphis albotecta'' *'' Graphis alpestris'' *'' Graphis analoga'' *''Graphis anfractuosa'' *''Graphis antillarum'' *'' Graphis aperiens'' *'' Graphis apertella'' *'' Graphis appendiculata'' *'' Graphis archeri'' *'' Graphis breussii'' *'' Graphis bulacana'' *'' Graphis caesiella'' *'' Graphis carassensis'' *'' Graphis caribica'' *'' Graphis centrifuga'' *'' Graphis cervina'' *'' Graphis chlorotica'' *''Graphis chrysocarpa'' *'' Graphis cincta'' *'' Graphis cinerea'' *''Graphis componens'' *''Graphis conturbata'' *''Graphis crassilabra'' *'' Graphis crebra'' *''Graphis crystallifera'' *''Graphis cycasicola'' *'' Graphis dealbata'' *''Graphis dendrogramma'' *''Graphis descissa'' *''Graphis disserpens'' *'' Graphis duplicata'' *'' Graphis dussii'' *'' Graphis eimeoensis'' *''Graphis elegans'' *'' Graphis elixiana'' *''Graphis emersa'' *''Graphis epimelaeana'' *'' Gr ...
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Thin-layer Chromatography
Thin-layer chromatography (TLC) is a chromatography technique used to separate non-volatile mixtures. Thin-layer chromatography is performed on a sheet of an inert substrate such as glass, plastic, or aluminium foil, which is coated with a thin layer of adsorbent material, usually silica gel, aluminium oxide (alumina), or cellulose. This layer of adsorbent is known as the stationary phase. After the sample has been applied on the plate, a solvent or solvent mixture (known as the mobile phase) is drawn up the plate via capillary action. Because different analytes ascend the TLC plate at different rates, separation is achieved. It may be performed on the analytical scale as a means of monitoring the progress of a reaction, or on the preparative scale to purify small amounts of a compound. TLC is an analytical tool widely used because of its simplicity, relative low cost, high sensitivity, and speed of separation. TLC functions on the same principle as all chromatography: a com ...
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Secondary Metabolite
Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the normal growth, development, or reproduction of the organism. Instead, they generally mediate ecological interactions, which may produce a selective advantage for the organism by increasing its survivability or fecundity. Specific secondary metabolites are often restricted to a narrow set of species within a phylogenetic group. Secondary metabolites often play an important role in plant defense against herbivory and other interspecies defenses. Humans use secondary metabolites as medicines, flavourings, pigments, and recreational drugs. The term secondary metabolite was first coined by Albrecht Kossel, a 1910 Nobel Prize laureate for medicine and physiology in 1910. 30 years later a Polish botanist Friedrich Czapek described secondary metabolit ...
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Septum
In biology, a septum (Latin for ''something that encloses''; plural septa) is a wall, dividing a cavity or structure into smaller ones. A cavity or structure divided in this way may be referred to as septate. Examples Human anatomy * Interatrial septum, the wall of tissue that is a sectional part of the left and right atria of the heart * Interventricular septum, the wall separating the left and right ventricles of the heart * Lingual septum, a vertical layer of fibrous tissue that separates the halves of the tongue. *Nasal septum: the cartilage wall separating the nostrils of the nose * Alveolar septum: the thin wall which separates the alveoli from each other in the lungs * Orbital septum, a palpebral ligament in the upper and lower eyelids * Septum pellucidum or septum lucidum, a thin structure separating two fluid pockets in the brain * Uterine septum, a malformation of the uterus * Vaginal septum, a lateral or transverse partition inside the vagina * Intermuscular sep ...
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Ascus
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Paraphyses
Paraphyses are erect sterile filament-like support structures occurring among the reproductive apparatuses of fungi, ferns, bryophytes and some thallophytes. The singular form of the word is paraphysis. In certain fungi, they are part of the fertile spore-bearing layer. More specifically, paraphyses are sterile filamentous hyphal end cells composing part of the hymenium of Ascomycota and Basidiomycota interspersed among either the asci or basidia respectively, and not sufficiently differentiated to be called cystidia A cystidium (plural cystidia) is a relatively large cell found on the sporocarp of a basidiomycete (for example, on the surface of a mushroom gill), often between clusters of basidia. Since cystidia have highly varied and distinct shapes that ar ..., which are specialized, swollen, often protruding cells. The tips of paraphyses may contain the pigments which colour the hymenium. In ferns and mosses, they are filament-like structures that are found on sporangia ...
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Hymenium
The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia (basidiomycetes) or paraphyses (ascomycetes). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive hyphae from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore. The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic Basidiomycota and Ascomycota. * In agarics, the hymenium is on the vertical faces of the gills. * In boletes and polypores, it is in a spongy mass of downward-pointing tubes. * In puffballs, ...
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Apothecia
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks of ...
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