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Grandinasus
''Esmeraldina'' is an extinct genus of holmiid trilobites. It lived during the late Atdabanian stage, which lasted from 530 to 524 million years ago during the early part of the Cambrian Period The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million .... References Cambrian trilobites Olenelloidea {{Redlichiida-stub ...
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Holmiidae
Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica (Scandinavia and the eastern seaboard of the Baltic Sea) and Western Laurentia (in the Great Basin of the US and northwestern Canada), and also in Morocco. Taxonomy Hupé (1953) defined the Holmiidae as a subfamily (Holmiinae) within the Olenellidae containing ''Holmia'', ''Kjerulfia'' and ''Bondonella''. Harrington et al. (1959) excluded ''Kjerulfia'', while assigning ''Schmidtiellus'' to it. Bergström (1973) included ''Holmia'', '' Elliptocephala'', ''Esmeraldina'', ''Schmidtiellus'' and '' Wanneria''. Repina (1979) assigned ''Holmia'', ''Schmidtiellus'', ''Kjerulfia'', ''Elliptocephala'', ''Bondonella'', ''Andalusiana'' and ''Holmiella'' to the Holmiinae, that together with the monotypic Callaviinae comprised the Hol ...
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Holmiidae
Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica (Scandinavia and the eastern seaboard of the Baltic Sea) and Western Laurentia (in the Great Basin of the US and northwestern Canada), and also in Morocco. Taxonomy Hupé (1953) defined the Holmiidae as a subfamily (Holmiinae) within the Olenellidae containing ''Holmia'', ''Kjerulfia'' and ''Bondonella''. Harrington et al. (1959) excluded ''Kjerulfia'', while assigning ''Schmidtiellus'' to it. Bergström (1973) included ''Holmia'', '' Elliptocephala'', ''Esmeraldina'', ''Schmidtiellus'' and '' Wanneria''. Repina (1979) assigned ''Holmia'', ''Schmidtiellus'', ''Kjerulfia'', ''Elliptocephala'', ''Bondonella'', ''Andalusiana'' and ''Holmiella'' to the Holmiinae, that together with the monotypic Callaviinae comprised the Hol ...
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Atdabanian
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera ''Olenellus'' or '' Redlichia'' around million years ago. Naming The International Commission on Stratigraphy has not officially named the 3rd stage of the Cambrian. The stage approximately corresponds to the "Atdabanian", which is used by geologists working in Siberia. Biostratigraphy The oldest trilobite known is ''Lemdadella'' which appears at the beginning of the '' Fallotaspis'' zone. The Cambrian radiatio ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinode ...
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Arthropoda
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Trilobita
Trilobites (; meaning "three lobes") are extinction, extinct marine arthropods that form the class (biology), class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian Stage (geology), stage of the Early Cambrian Period (geology), period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in Permian–Triassic extinction event, the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had ...
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Redlichiida
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are '' Fallotaspis'' (suborder Olenellina), and ''Lemdadella'' (suborder Redlichiina), both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian. Description Most redlichiids are rather flat (or have low dorso-ventral convexity) and their exoskeleton typically has an oval outline, ...
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Olenellina
Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. The earliest trilobites in the fossil record are arguably Olenellina, although the earliest Redlichiina and Eodiscina follow quickly. The suborder died out when the Lower passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines (or sutures) in the headshield, which in other trilobites assist the periodic moulting (or ecdysis), associated with arthropod growth. Some derived trilobites have lost facial sutures again (some Eodiscina, all Agnostina, and a few Phacopina), but all of these are blind, while all Olenellina have eyes. Taxonomy The suborder contains four superfamilies: Olenelloidea (with 3 families and 5 stemgroup genera), Judomioidea (with 1 family and 3 stemgroup genera), Nevadioidea, and Fallotaspidoidea (with 3 familie ...
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Olenelloidea
The Olenelloidea are a superfamily of trilobites, a group of extinct marine arthropods. They lived during the late Lower Cambrian and species occurred on all paleocontinents. Taxonomy Palmer and Repina assigned '' Gabriellus'' and ''Callavia'' to the Olenelloidea, but the ocular lobes in these two genera circumscribe the entire lateral border of the frontal lobe of the glabella, and thus, do not belong to this superfamily, but to the Judomioidea. Geyer assigned '' Cambropallas'' to the Olenelloidea, but it lacks the diagnostic features and appears to be closely related to '' Andalusiana'', and is now considered to be an advanced member of the Nevadioidea. Distribution In western Laurentia Fallotaspididae and Archaeaspididae proceed the first Olenelloidea. The Holmiidae occur at the late Atdabanian, and are contemporary with the Nevadioidea. They are followed by Biceratopsidae and Olenellidae. Description As with most early trilobites, the Olenelloidea have an almost ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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