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Gosslingiales
Gosslingiales is an order of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in various ways. Hao and Xue in 2013 used the presence or absence of terminal sporangia as a major dividing feature. The order Zosterophyllales was used for species with terminal as well as lateral sporangia, which were considered to have determinate growth, with their sporangia generally being arranged in spikes. The paraphyletic order Gosslingiales was used for species without terminal sporangia (i.e. with only lateral sporangia), which were considered to have indeterminate growth, with fertile branches generally circinate (initially curled up). Species assignable to the Gosslingiales made up about 9% of all confirmed species in the Early Devonian flora. Taxonomy Classification Hao and Xue consider the order Gosslingiales to be one of the two major divisions of the zost ...
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Zosterophyllum
''Zosterophyllum'' was a genus of Silurian-Devonian vascular land plants with naked branching axes on which usually kidney-shaped sporangia were arranged in lateral positions. It is the type genus for the group known as zosterophylls, thought to be part of the lineage from which modern lycophytes evolved. More than 20 species have been described. Description The diagnostic features of the genus have changed since its first description in 1892, as the original species (''Zosterophyllum myretonianum'') has become better known, and as other species have been discovered. ''Zosterophyllum'' is a vascular plant. The axes (stems) are naked, lacking leaves or outgrowths ("enations"). When branching occurs, the branches are either isotomous (equally sized) or pseudomonopodial (one branch is larger than the other but still clearly involves division of the original axis rather a distinct side growth). The sporangia are upright on short stalks. In face view, they are flattened, usually kid ...
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Sawdoniales
The Sawdoniales are an order or plesion of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in various ways. In their major cladistic study of early land plants, Kenrick and Crane placed most of the zosterophylls in the Sawdoniales (which they treated as a plesion). Like other zosterophylls, members of the Sawdoniales bore lateral, reniform sporangia. They branched dichotomously, and grew at the ends by unrolling (circinate vernation). Some had smooth stems, others were covered in small spines; fungal bodies have been reported in some spines. Taxonomy In 1997, Kenrick and Crane placed most of the zosterophylls in the plesion Sawdoniales, characterizing the group as having "marked bilateral symmetry". Their summary cladogram did not resolve the taxa within the Sawdoniales, other than placing '' Zosterophyllum divaricatum'' within the zosterophylls bu ...
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Discalis
''Discalis'' is a genus of extinct vascular plants of the Early Devonian (Pragian or Siegenian stage, around ). The name is derived from the Greek , referring to the disc-shaped sporangia (spore-forming organs). The genus was first described by Hao in 1989 based on fossil specimens from the Posongchong Formation, Wenshan district, Yunnan, China. Description The leafless sporophyte of ''D. longistipa'' consisted of creeping stems (axes) up to in diameter with many K- or H-shaped branches as well as upturned or trailing stems, slightly smaller in diameter, which also branched. All stems had irregularly arranged multicellular spines up to long with expanded tips. The stems which did not bear sporangia had coiled (circinnate) tips. Fertile stems bore disc-shaped sporangia laterally on stalks up to 5 mm long, forming open spikes. The sporangia, which were about in diameter, had spines like the stems, and split (dehisced) along their margin to release the trilete spores, whi ...
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Gosslingia
''Gosslingia'' was a genus of Early Devonian land plant with branching axes. Fossils have been from the Lochkovian to the Pragian, . A cladogram published in 2004 by Crane et al. places ''Gosslingia'' in the core of a paraphyletic stem group of broadly defined "zosterophylls", basal to the lycopsids (living and extinct clubmosses and relatives). Hao and Xue in 2013 used the absence of terminal sporangia to place the genus in the family Gosslingiaceae in the paraphyletic order Gosslingiales, a group considered to have indeterminate growth, with fertile branches generally showing circinate vernation (initially curled up). Kenrick and Crane in 1997 also placed the genus in the family Gosslingiaceae, but they place this family in the order Sawdoniales The Sawdoniales are an order or plesion of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in ...
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Sawdonia
Sawdonia is an extinct genus of early vascular plants, known from the Upper Silurian to the Lower Carboniferous (). ''Sawdonia'' is best recognized by the large number of spikes (enations) covering the plant. These are vascular plants that do not have vascular systems in their enations. The first species of this genus (''Sawdonia ornata'') was described in 1859 by Sir J. William Dawson and, was originally attributed to the genus '' Psilophyton''. He named this plant ''Psilophyton princeps''. In 1971 Francis Hueber proposed a new genus for this species due to its "Divergent technical characters from the generic description for ''Psilophyton''." The holotype used for description is Dawson Collection Number 48, pro parte, Museum Specimen Number 3243. (See Dawson 1871, Plate IX, fig 101.) Sir J. William Dawson Collection, Peter Redpath Museum, McGill University, Montreal, Quebec, Canada. Morphology These plants are described by Hueber as having monopodially branched stems, that ar ...
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Konioria
''Konioria'' was a genus of early land plant with branching axes. Known fossils are of Early Devonian age (). A cladogram published in 2004 by Crane et al. places ''Konioria'' in the core of a paraphyletic stem group of broadly defined "zosterophylls", basal to the lycopsids (living and extinct clubmosses and relatives). Hao and Xue in 2013 used the absence of terminal sporangia to place the genus in the paraphyletic order Gosslingiales, a group considered to have indeterminate growth, with fertile branches generally showing circinate vernation Vernation (from ''vernal'' meaning ''spring'', since that is when leaves spring forth in temperate regions) is the formation of new leaves or fronds. In plant anatomy, it is the arrangement of leaves in a bud. In pine species, new leaves are short ... (initially curled up). References External links Cladogramfrom Zosterophylls Prehistoric lycophyte genera Devonian plants Lochkovian {{paleo-lycophyte-stub ...
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Oricilla
''Oricilla'' was a genus of Early Devonian land plant with branching axes. Fossils have been found from the Pragian to the Emsian (). A cladogram published in 2004 by Crane et al. places ''Oricilla'' in the core of a paraphyletic stem group of broadly defined "zosterophylls", basal to the lycopsids (living and extinct clubmosses and relatives). Hao and Xue in 2013 used the absence of terminal sporangia to place the genus in the paraphyletic order Gosslingiales, a group of zosterophylls considered to have indeterminate growth, with fertile branches generally showing circinate vernation (initially curled up). Kenrick and Crane in 1997 also placed the genus in the family Gosslingiaceae, but they place this family in the order Sawdoniales The Sawdoniales are an order or plesion of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in various ways. In ...
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Gosslingiaceae
Gosslingiaceae is a family of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and are considered to share an ancestor with the living lycophytes. The family is variously placed in the order Sawdoniales or the order Gosslingiales Gosslingiales is an order of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in various ways. Hao and Xue in 20 .... Genera Genera that have been placed in this family by Kenrick and Crane in 1997 and Hao and Xue in 2013 are shown in the table below. References Zosterophylls Prehistoric plant families {{paleobotany-stub ...
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Crenaticaulis
''Crenaticaulis'' was an early genus of slender, dichotomously branching, leafless land plants, known from the Devonian period and first described in 1969. They were probably allied to the zosterophylls, and are assigned to subdivision Zosterophyllophytina, or class Zosterophyllopsida. They bore branches and scalariform tracheids. A cladogram published in 2004 by Crane et al. places ''Crenaticaulis'' in the core of a paraphyletic stem group of broadly defined "zosterophylls", basal to the lycopsids (living and extinct clubmosses and relatives). Hao and Xue in 2013 used the absence of terminal sporangia to place the genus in the family Gosslingiaceae in the paraphyletic order Gosslingiales, a group considered to have indeterminate growth, with fertile branches generally showing circinate vernation (initially curled up). Kenrick and Crane in 1997 placed the genus in the family Sawdoniaceae in the order Sawdoniales The Sawdoniales are an order or plesion of extinct zosterophyl ...
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Sawdoniaceae
Sawdoniaceae is a family of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and are considered to share an ancestor with the living lycophytes. The family is recognized by some sources, and placed in the Sawdoniales The Sawdoniales are an order or plesion of extinct zosterophylls. The zosterophylls were among the first vascular plants in the fossil record, and share an ancestor with the living lycophytes. The group has been divided up in various ways. In t .... Other sources do not recognize the family, and place some of its members in the family Gosslingiaceae. Genera Genera that have been placed in this family by Kenrick and Crane in 1997 are shown in the table below, along with their treatment by Hao and Xue in 2013. References Zosterophylls Prehistoric plant families {{paleobotany-stub ...
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