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Glyptosaurus
Glyptosaurinae is an extinct subfamily of anguid lizards that lived in the Northern Hemisphere during the Late Cretaceous and the Paleogene. Description Glyptosaurines are known primarily from their osteoderms, scale-like pieces of bone that are embedded in the skin and cover much of their bodies. The shape and extent of the osteoderms in glyptosaurines are similar to those seen in an unrelated group of lizards called Monstersauria, which includes the living Gila monster and beaded lizard. The osteoderms of glyptosaurines are unusually complex, consisting of four distinct layers of bony tissue. These tissues may have derived from both the dermis (the lower layer of the skin) and the epidermis (the outer layer of the skin) during their development in the embryo. The tissue forming the outermost layer of glyptosaurine osteoderms is similar to tooth enamel and has even been given its own name, osteodermine. Glyptosaurines have been split into the subgroups Melanosaurini and Glyptos ...
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Glyptosaurus
Glyptosaurinae is an extinct subfamily of anguid lizards that lived in the Northern Hemisphere during the Late Cretaceous and the Paleogene. Description Glyptosaurines are known primarily from their osteoderms, scale-like pieces of bone that are embedded in the skin and cover much of their bodies. The shape and extent of the osteoderms in glyptosaurines are similar to those seen in an unrelated group of lizards called Monstersauria, which includes the living Gila monster and beaded lizard. The osteoderms of glyptosaurines are unusually complex, consisting of four distinct layers of bony tissue. These tissues may have derived from both the dermis (the lower layer of the skin) and the epidermis (the outer layer of the skin) during their development in the embryo. The tissue forming the outermost layer of glyptosaurine osteoderms is similar to tooth enamel and has even been given its own name, osteodermine. Glyptosaurines have been split into the subgroups Melanosaurini and Glyptos ...
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Stenoplacosaurus
''Stenoplacosaurus'' is an extinct genus of glyptosaurine anguid lizards from the Eocene of Inner Mongolia, China. The genus is monotypic, containing only the species ''Stenoplacosaurus mongoliensis''. Taxonomy ''Stenoplacosaurus'' was originally named ''Helodermoides mongoliensis'' by Sullivan (1979) for remains found in the Shara Murun Formation of Inner Mongolia Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an autonomous region of the People's Republic of China. Its border includes most of the length of China's border with the country of Mongolia. Inner Mongolia also accounts for a ..., China. In their redescription of '' Placosaurus'', Sullivan and Auge (2006) transferred ''H. mongoliensis'' to ''Placosaurus'', as ''P. mongoliensis''. However, Sullivan and Dong (2018) found it to be generically distinct from the ''Placosaurus'' type species and renamed it ''Stenoplacosaurus''. References {{Extinct squamates, A. Anguids Eocene reptiles of Asia ...
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Helodermoides Tuberculatus
''Helodermoides'' is an extinct genus of anguid lizards from the Oligocene of North America. The genus is monotypic, including only the species ''Helodermoides tuberculatus''. ''Helodermoides'' belongs to an extinct subfamily of anguids called Glyptosaurinae. In addition to many fragmentary bones, several complete skeletons of ''Helodermoides'' are known. Like other glyptosaurines, ''Helodermoides'' was covered in small scale-like bones called osteoderms. The osteoderms covering its skull are hexagonal, tightly interlocking, raised, and rounded. One fossil of ''Helodermoides'' preserves a fused mass of osteoderms at the tip of a shortened tail, thought to represent healing after the end of the tail fell off. The tail would not have been able to grow back because the osteoderms formed a thick bony cap preventing growth. The ability to lose a tail, called autotomy, is also present in living anguid Anguidae refers to a large and diverse family of lizards native to the Northern Hem ...
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Northern Hemisphere
The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's North Pole. Owing to Earth's axial tilt of 23.439281°, winter in the Northern Hemisphere lasts from the December solstice (typically December 21 UTC) to the March equinox (typically March 20 UTC), while summer lasts from the June solstice through to the September equinox (typically on 23 September UTC). The dates vary each year due to the difference between the calendar year and the astronomical year. Within the Northern Hemisphere, oceanic currents can change the weather patterns that affect many factors within the north coast. Such events include El Niño–Southern Oscillation. Trade winds blow from east to west just above the equator. The winds pull surface water with them, creating currents, which flow westward due to the Coriolis e ...
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Evolutionary Grade
A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit. Definition An evolutionary grade is a group of species united by morphological or physiological traits, that has given rise to another group that has major differences from the ancestral condition, and is thus not considered part of the ancestral group, while still having enough similarities that we can group them under the same clade. The ancestral group will not be phylogenetically complete (i.e. will not form a clade), so will represent a paraphyletic taxon. In order to fully understand evolutionary grades, one must first get a better understanding of Phylogenetics, defined as "''In biology, is the study of the evolutionary history and relationships among individuals or groups of organisms (e.g. species, or populations). These relationships are discovered through phylogenetic i ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Tooth Enamel
Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The other major tissues are dentin, cementum, and Pulp (tooth), dental pulp. It is a very hard, white to off-white, highly mineralised substance that acts as a barrier to protect the tooth but can become susceptible to degradation, especially by acids from food and drink. Calcium hardens the tooth enamel. In rare circumstances enamel fails to form, leaving the underlying dentin exposed on the surface. Features Enamel is the hardest substance in the human body and contains the highest percentage of minerals (at 96%),Ross ''et al.'', p. 485 with water and organic material composing the rest.Ten Cate's Oral Histology, Nancy, Elsevier, pp. 70–94 The primary mineral is hydroxyapatite, which is a crystalline calcium phosphate. Enamel is formed o ...
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Epidermis (skin)
The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water released from the body into the atmosphere through transepidermal water loss. The epidermis is composed of multiple layers of flattened cells that overlie a base layer (stratum basale) composed of columnar cells arranged perpendicularly. The layers of cells develop from stem cells in the basal layer. The human epidermis is a familiar example of epithelium, particularly a stratified squamous epithelium. The word epidermis is derived through Latin , itself and . Something related to or part of the epidermis is termed epidermal. Structure Cellular components The epidermis primarily consists of keratinocytes ( proliferating basal and differentiated suprabasal), which comprise 90% of its cells, but also contains melanocytes, Langerhans c ...
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Dermis
The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided into two layers, the superficial area adjacent to the epidermis called the papillary region and a deep thicker area known as the reticular dermis.James, William; Berger, Timothy; Elston, Dirk (2005). ''Andrews' Diseases of the Skin: Clinical Dermatology'' (10th ed.). Saunders. Pages 1, 11–12. . The dermis is tightly connected to the epidermis through a basement membrane. Structural components of the dermis are collagen, elastic fibers, and extrafibrillar matrix.Marks, James G; Miller, Jeffery (2006). ''Lookingbill and Marks' Principles of Dermatology'' (4th ed.). Elsevier Inc. Page 8–9. . It also contains mechanoreceptors that provide the sense of touch and thermoreceptors that provide the sense of heat. In addition, hair follicles, ...
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Beaded Lizard
The Mexican beaded lizard (''Heloderma horridum'') is a species of lizard in the Family (biology), family Helodermatidae, one of the two species of venomous beaded lizards found principally in Mexico and southern Guatemala. It and congener (biology), the other member of the same genus, the Gila monster (''Heloderma suspectum''), are the only lizards known to have evolved an overt venom delivery system. The Mexican beaded lizard is larger than the Gila monster, with duller coloration, black with yellowish bands. As it is a specialized predator that feeds primarily upon eggs, the primary use of its venom is still a source of debate among scientists. This venom has been found to contain several enzymes useful for manufacturing drugs in the treatment of diabetes, and research on the pharmacological use of its venom is ongoing. Threatened throughout its range by overcollection and Habitat destruction, habitat loss, it is a CITES protected species. The Heloderma charlesbogerti, Guatem ...
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Gila Monster
The Gila monster (''Heloderma suspectum'', ) is a species of venomous lizard native to the Southwestern United States and the northwestern Mexican state of Sonora. It is a heavy, typically slow-moving reptile, up to long, and it is the only venomous lizard native to the United States. Its venomous close relatives, the four beaded lizards (all former subspecies of ''Heloderma horridum'') inhabit Mexico and Guatemala. The Gila monster is sluggish in nature, so it is not generally dangerous and very rarely poses a real threat to humans. However, it has a fearsome reputation and is sometimes killed in spite of the species being protected by state law in Arizona. History The name "Gila" refers to the Gila River Basin in the U.S. states of Arizona and New Mexico, where the Gila monster was once plentiful. ''Heloderma'' means "studded skin", from the Ancient Greek words (), "the head of a nail or stud", and (), "skin". ''Suspectum'' comes from the describer, paleontologist Edwa ...
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