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Gladocephaloideus Anatomy
''Gladocephaloideus'' is a genus of ctenochasmatid ctenochasmatoid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now western Liaoning, China. Discovery ''Gladocephaloideus'' is known from a complete skull and partial postcranial skeleton including traces of hair-like pycnofibres. It was collected from the Jingangshan Bed of the Yixian Formation, dating to the early Aptian stage, about 121 million years ago. It was named by Lü Junchang, Ji Qiang, Wei Xuefang and Liu Yongqing in 2012 and the type species is ''Gladocephaloideus jingangshanensis''. The generic name is derived from Latin ''gladius'', "sword", and Greek κεφαλή, ''kephalè'', "head", en εἶδος, ''eidos'', "form", referring to the elongated form of the skull. The specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Type Species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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Pterofiltrus
''Pterofiltrus'' is a genus of ctenochasmatid pterosaur from the Early Cretaceous of western Liaoning, China. Discovery and naming ''Pterofiltrus'' was named by Jiang Shunxing and Wang Xiaolin in 2011. The type species is ''Pterofiltrus qiui''. The generic name is derived from Greek πτερόν, ''pteron'', "wing", and a Mediaeval Latin ''filtrum'', "felt" or "filter", in reference to the dentition. The specific name honours Professor Qiu Zhanxiang. The holotype of ''Pterofiltrus'', IVPP V12339, was early 21st century discovered at Zhangjiagou in Liaoning Province in a layer of the Jianshangou Beds of the lower Yixian Formation dating from the early Aptian, about 125 million years old. It consists of a disarticulated skull compressed on a slab together with two rami of a hyoid and the first two neck vertebrae. Description The skull of ''Pterofiltrus'' is very elongated, with an estimated length of 208 millimetres. It has a smooth, slightly concave, upper profile lacking any b ...
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Sister Species
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "cha ...
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Gallodactylidae
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey. History Gallodactylidae was named to contain ''Gallodactylus'' (now usually considered a synonym of ''Cycnorhamphus'') and its closest relatives. Many subsequent studies, however, showed that ''Gallodactylus'' did not form a clade with any non-synonymous pterosaurs that were not themselves part of a different family, and so the name was often ignored. The name returned to common use with the discovery of ''Gladocephaloideus'', a Chinese pterosaur species that shared many similarities with ''Cycnorhamphus''. Among other features, the Gallodactylidae was distinguished by ...
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Gladocephaloideus Anatomy
''Gladocephaloideus'' is a genus of ctenochasmatid ctenochasmatoid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now western Liaoning, China. Discovery ''Gladocephaloideus'' is known from a complete skull and partial postcranial skeleton including traces of hair-like pycnofibres. It was collected from the Jingangshan Bed of the Yixian Formation, dating to the early Aptian stage, about 121 million years ago. It was named by Lü Junchang, Ji Qiang, Wei Xuefang and Liu Yongqing in 2012 and the type species is ''Gladocephaloideus jingangshanensis''. The generic name is derived from Latin ''gladius'', "sword", and Greek κεφαλή, ''kephalè'', "head", en εἶδος, ''eidos'', "form", referring to the elongated form of the skull. The specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set ...
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Metatarsal
The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the medial side (the side of the great toe): the first, second, third, fourth, and fifth metatarsal (often depicted with Roman numerals). The metatarsals are analogous to the metacarpal bones of the hand. The lengths of the metatarsal bones in humans are, in descending order, second, third, fourth, fifth, and first. Structure The five metatarsals are dorsal convex long bones consisting of a shaft or body, a base (proximally), and a head (distally).Platzer 2004, p. 220 The body is prismoid in form, tapers gradually from the tarsal to the phalangeal extremity, and is curved longitudinally, so as to be concave below, slightly convex above. The base or posterior extremity is wedge-shaped, articulating proximally with the tarsal bones, and by it ...
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Gladocephaloideus Jingangshanensis
''Gladocephaloideus'' is a genus of ctenochasmatid ctenochasmatoid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now western Liaoning, China. Discovery ''Gladocephaloideus'' is known from a complete skull and partial postcranial skeleton including traces of hair-like pycnofibres. It was collected from the Jingangshan Bed of the Yixian Formation, dating to the early Aptian stage, about 121 million years ago. It was named by Lü Junchang, Ji Qiang, Wei Xuefang and Liu Yongqing in 2012 and the type species is ''Gladocephaloideus jingangshanensis''. The generic name is derived from Latin ''gladius'', "sword", and Greek κεφαλή, ''kephalè'', "head", en εἶδος, ''eidos'', "form", referring to the elongated form of the skull. The specific name refers to the provenance from the Jiangangshan. The holotype, IG-CAGS-08-07, includes the almost complete skull, the lower jaws, the front neck vertebrae and part of a hindlimb. In 2016, a second speci ...
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Jiufotang Formation
The Jiufotang Formation (Chinese: 九佛堂组, pinyin: ''jiǔfótáng zǔ'') is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms (see Jehol Biota). It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with estimates ranging from the Late Jurassic to the Early Cretaceous. New uranium-lead dates reveal the formation is deposited in the Aptian stage of the Early Cretaceous. Fossils of ''Microraptor'' and ''Jeholornis'' are from the Jiufotang. Fossil content Choristoderans Fish Mammaliamorphs Several mammaliamorph specimens have been found from the Jiufotang, but only two have been formally described and named. Ornithischians Pterosaurs Saurischians A large titanosaur is present in the formation. Enantiornithines Euornithines Misc theropods See also * Yixian Formation * List of dinosaur-bearing rock for ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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