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Frullania Nisquallensis
''Frullania nisquallensis'', commonly known as hanging millipede liverwort, is a reddish-brown species of liverwort in the family Frullaniaceae. It is found in western Washington and British Columbia, including Vancouver Island. The plant grows in mats, sometimes in mats that hang from tree branches (particularly those of alders, or maples), or growing close to the substrate. The leaves are small (1 mm long) and flat, with the lower leaves being slightly smaller than those growing farther up the stem.''Plants of the Pacific Northwest Coast''. Lone Pine Publishing, 1994. p. 444 Secondary metabolites This species contains the tridepside A depside is a type of polyphenolic compound composed of two or more monocyclic aromatic units linked by an ester bond. Depsides are most often found in lichens, but have also been isolated from higher plants, including species of the Ericaceae, ... compound tenuiron, and the sesquiterpenes (-)-frullanolide and costunolide, the latter ...
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Marchantiophyta
The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information. It is estimated that there are about 9000 species of liverworts. Some of the more familiar species grow as a flattened leafless thallus, but most species are leafy with a form very much like a flattened moss. Leafy species can be distinguished from the apparently similar mosses on the basis of a number of features, including their single-celled rhizoids. Leafy liverworts also differ from most (but not all) mosses in that their leaves never have a costa (present in many mosses) and may bear marginal cilia (very rare in mosses). Other differences are not universal for all mosses and liverworts, but the occurrence of leaves arranged in three ranks, the presence of deep lobes or segmented leaves, or a lack of clearly diff ...
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Frullaniaceae
''Frullania'' is the only genus of liverworts in family Frullaniaceae. It contains the following species: A *''Frullania aculeata'' Taylor, 1846 *''Frullania acutata'' Caspary, 1887 *''Frullania acutiloba'' Gerola, 1947 *'' Frullania akiyamae'' Hattori, 1986 *'' Frullania albertii'' Stephani, 1916 *'' Frullania allanii'' Hodgson, 1949 *'' Frullania allionii'' Stephani, 1910 *'' Frullania alpina'' Stephani, 1911 *''Frullania alstonii'' Verdoorn, 1930 *''Frullania alstonii'' var. ''pfleidereri'' Hattori, 1972 *'' Frullania alternans'' Nees In G., L. & N., 1845 *''Frullania amamiensis'' Kamimura, 1968 *''Frullania ambronnii'' Stephani, 1916 *''Frullania amplicrania'' Stephani, 1910 *''Frullania ampullifera'' Jack & Stephani In Stephani, 1894 *'' Frullania anderssonii'' Ångström, 1873 *'' Frullania angulata'' Mitten, 1863 *''Frullania angulata'' F. ''Serratoides'' Vanden Berghen, 1983 *''Frullania angulata'' var. ''laciniata'' Demaret & Vanden Berghen, 1950 *'' Frullania angustis ...
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Washington (U
Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered on Washington, D.C. * George Washington (1732–1799), the first president of the United States Washington may also refer to: Places England * Washington, Tyne and Wear, a town in the City of Sunderland metropolitan borough ** Washington Old Hall, ancestral home of the family of George Washington * Washington, West Sussex, a village and civil parish Greenland * Cape Washington, Greenland * Washington Land Philippines *New Washington, Aklan, a municipality *Washington, a barangay in Catarman, Northern Samar *Washington, a barangay in Escalante, Negros Occidental *Washington, a barangay in San Jacinto, Masbate *Washington, a barangay in Surigao City United States * Washington, Wisconsin (other) * Fort Washington (disambiguati ...
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British Columbia
British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, forests, lakes, mountains, inland deserts and grassy plains, and borders the province of Alberta to the east and the Yukon and Northwest Territories to the north. With an estimated population of 5.3million as of 2022, it is Canada's third-most populous province. The capital of British Columbia is Victoria and its largest city is Vancouver. Vancouver is the third-largest metropolitan area in Canada; the 2021 census recorded 2.6million people in Metro Vancouver. The first known human inhabitants of the area settled in British Columbia at least 10,000 years ago. Such groups include the Coast Salish, Tsilhqotʼin, and Haida peoples, among many others. One of the earliest British settlements in the area was Fort Victoria, established ...
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Vancouver Island
Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian Provinces and territories of Canada, province of British Columbia. The island is in length, in width at its widest point, and in total area, while are of land. The island is the largest by area and the most populous along the west coasts of the Americas. The southern part of Vancouver Island and some of the nearby Gulf Islands are the only parts of British Columbia or Western Canada to lie south of the 49th parallel north, 49th parallel. This area has one of the warmest climates in Canada, and since the mid-1990s has been mild enough in a few areas to grow Mediterranean Sea, Mediterranean crops such as olives and lemons. The population of Vancouver Island was 864,864 as of 2021. Nearly half of that population (~400,000) live in the metropolitan area of Greater Victoria, the capital city of British Columbia. Other notable cities and towns on Vancouver Island include Nanaimo, Port Alberni, ...
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Tridepside
A depside is a type of polyphenolic compound composed of two or more monocyclic aromatic units linked by an ester bond. Depsides are most often found in lichens, but have also been isolated from higher plants, including species of the Ericaceae, Lamiaceae, Papaveraceae and Myrtaceae. Certain depsides have antibiotic, anti-HIV, antioxidant, and anti-proliferative activity ''in vitro''. As inhibitors of prostaglandin synthesis and leukotriene B4 biosynthesis, some depsides have ''in vitro'' anti-inflammatory activity. A depsidase is a type of enzyme that cuts depside bonds. One such enzyme is tannase. Examples Gyrophoric acid, found in the lichen ''Cryptothecia rubrocincta'', is a depside. Merochlorophaeic acid, isolated from lichens of the genus ''Cladonia'', is an inhibitor of prostaglandin synthesis. Some depsides are described as anti-HIV. See also * Salsalate homodimer formed from self-condensation of salicylic acid Salicylic acid is an organic compound with the form ...
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Sesquiterpene
Sesquiterpenes are a class of terpenes that consist of three isoprene units and often have the molecular formula C15H24. Like monoterpenes, sesquiterpenes may be cyclic or contain rings, including many unique combinations. Biochemical modifications such as oxidation or rearrangement produce the related sesquiterpenoids. Sesquiterpenes are found naturally in plants and insects, as semiochemicals, e.g. defensive agents or pheromones. Biosynthesis and examples The reaction of geranyl pyrophosphate with isopentenyl pyrophosphate results in the 15-carbon farnesyl pyrophosphate (FPP), which is an intermediate in the biosynthesis of sesquiterpenes such as farnesene. Cyclic sesquiterpenes are more common than cyclic monoterpenes because of the increased chain length and additional double bond in the sesquiterpene precursors. In addition to common six-membered ring systems such as the ones found in zingiberene and bisacurone, cyclization of one end of the chain to the other end can l ...
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