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Frond Dimorphism
Frond dimorphism refers to a difference in ferns between the fertile and sterile fronds. Since ferns, unlike flowering plants, bear spores on the leaf blade itself, this may affect the form of the frond itself. In some species of ferns, there is virtually no difference between the fertile and sterile fronds, such as in the genus ''Dryopteris'', other than the mere presence of the sori, or fruit-dots, on the back of the fronds. Some other species, such as ''Polystichum acrostichoides'' (Christmas fern), or some ferns of the genus ''Osmunda'', feature dimorphism on a portion of the frond only. Others, such as some species of ''Blechnum'' and ''Woodwardia'', have fertile fronds that are markedly taller than the sterile. Still others, such as ''Osmunda cinnamomea'' (Cinnamon fern), or plants of the family Onocleaceae, have fertile fronds that are completely different from the sterile. Only members of the ''Onocleaceae'' and ''Blechnaceae'' exhibit a propensity towards dimorphy, while ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Osmunda Cinnamomea
''Osmundastrum'' is genus of leptosporangiate ferns in the family Osmundaceae with one living species, ''Osmundastrum cinnamomeum'', the cinnamon fern. It is native to the Americas and eastern Asia, growing in swamps, bogs and moist woodlands. In North America it occurs from southern Labrador west to Ontario, and south through the eastern United States to eastern Mexico and the West Indies; in South America it occurs west to Peru and south to Paraguay. In Asia it occurs from southeastern Siberia south through Japan, Korea, China and Taiwan to Myanmar, Thailand and Vietnam. ''Osmundastrum cinnamomeum'' has a fossil record extending into the Late Cretaceous of North America, approximately 70 million years ago, making it one of the oldest living plant species. The fossil records of the genus extend into the Triassic. Characteristics ''Osmundastrum cinnamomeum'' is a deciduous herbaceous plant that produces separate fertile and sterile fronds. The sterile fronds are spreading, ...
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Springer Publishing
Springer Publishing Company is an American publishing company of academic journals and books, focusing on the fields of nursing, gerontology, psychology, social work, counseling, public health, and rehabilitation (neuropsychology). It was established in 1951 by Bernhard Springer, a great-grandson of Julius Springer, and is based in Midtown Manhattan, New York City. History Springer Publishing Company was founded in 1950 by Bernhard Springer, the Berlin-born great-grandson of Julius Springer, who founded Springer-Verlag (now Springer Science+Business Media). Springer Publishing's first landmark publications included ''Livestock Health Encyclopedia'' by R. Seiden and the 1952 ''Handbook of Cardiology for Nurses''. The company's books soon branched into other fields, including medicine and psychology. Nursing publications grew rapidly in number, as Modell's ''Drugs in Current Use'', a small annual paperback, sold over 150,000 copies over several editions. Solomon Garb's ''Labor ...
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Waltham, Massachusetts
Waltham ( ) is a city in Middlesex County, Massachusetts, United States, and was an early center for the labor movement as well as a major contributor to the American Industrial Revolution. The original home of the Boston Manufacturing Company, the city was a prototype for 19th century industrial city planning, spawning what became known as the Waltham-Lowell system of labor and production. The city is now a center for research and higher education, home to Brandeis University and Bentley University as well as industrial powerhouse Raytheon Technologies. The population was 65,218 at the census in 2020. Waltham has been called "watch city" because of its association with the watch industry. Waltham Watch Company opened its factory in Waltham in 1854 and was the first company to make watches on an assembly line. It won the gold medal in 1876 at the Philadelphia Centennial Exposition. The company produced over 35 million watches, clocks and instruments before it closed in 1957. Histo ...
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Rolla M
Rolla is an Italian surname and also a diminutive for the Toyota Corolla. Rolla may refer to: People Surname *Alessandro Rolla (1757–1841), Italian composer, violin and viola virtuoso * Antonio Rolla (1798–1837), Italian composer, violin and viola virtuoso Given name * Rolla Anderson (born 1920), American former football and basketball player and coach *Rolla Daringer (1888–1974), American baseball shortstop *Rolla Dyer (1886–1971), American physician * Rolla Mapel (1890–1966), American baseball pitcher * Rolla C. McMillen (1880–1961), American politician from Illinois * Rolla Norman (1889–1971), French actor *Rolla Wells (1856–1944), American politician from Missouri Places ;United States * Rolla, Kansas *Rolla, Missouri **Missouri University of Science and Technology, formerly the University of Missouri–Rolla *Rolla, North Dakota ;Canada * Rolla, British Columbia ;Norway *Rolla (Troms), an island in Troms county, in the municipality of Ibestad ;United Arab ...
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Edwin Copeland
Edwin Bingham Copeland (September 30, 1873 – March 16, 1964) was an American botanist and agriculturist. He is known for founding the University of the Philippines College of Agriculture at Los Baños, Laguna and for being one of the America's leading pteridologists (one who studies ferns). Life In 1903, he and his family moved to the Philippines, where he worked as a Systematic Botanist for the Bureau of Science. Wagner, W.H. Jr. 1964Edwin Bingham Copeland (1873–1964) and his contributions to Pteridology American Fern Journal 54(4): 177–188. In 1909, he founded the University of the Philippines College of Agriculture at Los Baños, Laguna, now part of the University of the Philippines Los Baños, and served as its dean and also as a professor of plant physiology for eight years (1909–1917). In 1917, he returned to the United States and was a leading rice grower in Chico, California. In 1927, he began work as an Associate Curator at the University of California, Berk ...
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Thelypteridaceae
Thelypteridaceae is a family of about 900 species of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is placed in the suborder Aspleniineae. Alternatively, the family may be submerged in a very broadly defined family Aspleniaceae as the subfamily Thelypteridoideae. The ferns are terrestrial, with the exception of a few which are lithophytes (grow on rocks). The bulk of the species are tropical, although there are a number of temperate species. These ferns typically have creeping rhizomes. The fronds are simply pinnate to pinnate-pinnatifid. There is either no frond dimorphism or only mild dimorphism, either open venation or very simple anastomosing. The sori are mostly reniform in shape and have indusia, except for the ''Phegopteris'' group. Classification During the early and mid 1900's all thelypterioid ferns were included in the genus ''Dryopteris'' because of the sorus shape. However, there are a great many differ ...
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Athyriaceae
The Athyriaceae (ladyferns and allies) are a family of terrestrial ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treated as the subfamily Athyrioideae of a very broadly defined family Aspleniaceae. The family has with a cosmopolitan distribution. Description Species of the Athyriaceae are terrestrial or lithophytic, less commonly aquatic. They grow from various kinds of rhizome: short or long, creeping or erect, branched or not. The distribution and evolution of characters in the family is complex, and the genera have few constant features by which they can be identified. The sporangia have stalks two or three cells wide in the middle, and contain brown monolete spores. Taxonomy Earlier classifications The family was first created by Arthur H.G. Alston in 1956. It has had a varied history. In 2014, Christenhusz ...
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Blechnaceae
Blechnaceae is a family of ferns in the order Polypodiales, with a cosmopolitan distribution. Its status as a family and the number of genera included have both varied considerably. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family has 24 genera, and excludes genera placed in the separate family Onocleaceae. The family is divided into three subfamilies, including Blechnoideae s.s. Alternatively, the entire family may be treated as the subfamily Blechnoideae s.l. of a very broadly defined family Aspleniaceae, and include genera others place in Onocleaceae. Description Most are ground dwelling, some are climbers, such as ''Stenochlaena''. A characteristic feature of many species is that the young opening fronds are usually tinged with red. Taxonomy The family was created by Newman in 1844. In 2014, Christenhusz and Chase submerged it as subfamily Blechnoideae within the family Aspleniaceae and included Onocleaceae in it. The PPG I classifica ...
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Onocleaceae
Onocleaceae is a small family of terrestrial ferns in the order Polypodiales. It is placed in the suborder Aspleniineae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family, along with Blechnaceae, may be placed in a very broadly defined family Aspleniaceae as the subfamily Blechnoideae. The family may contain from one to four genera, consisting of five species largely in north temperate climes. The four genera, '' Matteuccia'', ''Onoclea'', '' Onocleopsis'' and '' Pentarhizidium'', may be included under the single genus ''Onoclea''. Description Members of the family have the following characteristics, being distinguished by having strongly dimorphic fronds, with the fertile fronds different from the sterile fronds. The rhizomes are long- to short-creeping to ascending, and sometimes stoloniferous (''Matteuccia'' and ''Onocleopsis''). The leaves are strongly dimorphic and the petioles have two vascular bundles uniting distally into a gut ...
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Woodwardia
''Woodwardia'' is a genus of ferns in the family Blechnaceae, in the suborder Aspleniineae (eupolypods II) of the order Polypodiales. Species are known as chain ferns. The genus is native to warm temperate and subtropical regions of the Northern Hemisphere. They are large ferns, with fronds growing to 50–300 cm long depending on the species. The fossil record of the genus extends to the Paleocene. Taxonomy ''Woodwardia'' was first described by James Edward Smith in 1793. It was named after Thomas Jenkinson Woodward. When broadly circumscribed, the genus contains about 15 species (plus some hybrids). In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the genera '' Anchistea'' and '' Lorinseria'' (each with one species) are kept separate. Species , Plants of the World Online accepts the following species, excluding those placed in other genera in the PPG I system. *'' Woodwardia auriculata'' Blume *'' Woodwardia fimbriata'' Sm. *'' Woodwardia harlandii'' H ...
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Frond
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the large leaves of cycads, as well as palms (Arecaceae) and various other flowering plants, such as mimosa or sumac. "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns and algae it may be applied to smaller and undivided leaves. Fronds have particular terms describing their components. Like all leaves, fronds usually have a stalk connecting them to the main stem. In botany, this leaf stalk is generally called a petiole, but in regard to fronds specifically it is called a stipe, and it supports a flattened blade (which may be called a lamina), and the continuation of the stipe into this portion is called the rachis. The blades may be simple (undivided), pinnatifid ( ...
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