Euichthyosauria
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Euichthyosauria
Merriamosauria is an extinct clade of ichthyosaurs. It was named by Ryosuke Motani in his 1999 analysis of the relationships of ichthyopterygian marine reptiles and was defined in phylogenetic terms as a stem-based taxon including "the last common ancestor of ''Shastasaurus pacificus'' and '' Ichthyosaurus communis'', and all of its descendants." The name honours John Campbell Merriam. Based on this definition, Merriamosauria includes most ichthyosaurs except for several Triassic groups such as the clade Mixosauria, the family Cymbospondylidae, and perhaps the family Toretocnemidae. Merriamosaurs are characterized by features in their pectoral girdles and limb bones, including an extensive connection between the scapula and the coracoid bone, the absence of the first metacarpal and the absence of a pisiform bone. In addition to ''Shastasaurus,'' this clade includes ''Shonisaurus, Guizhouichthyosaurus ''Guizhouichthyosaurus'' is an extinct genus of ichthyosaur which is kn ...
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Ichthyosaur
Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, although the term is now used more for the parent clade of the Ichthyosauria). Ichthyosaurs thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago ( Ma) and at least one species survived until about 90 million years ago, into the Late Cretaceous. During the Early Triassic epoch, ichthyosaurs and other ichthyosauromorphs evolved from a group of unidentified land reptiles that returned to the sea, in a development similar to how the mammalian land-dwelling ancestors of modern-day dolphins and whales returned to the sea millions of years later, which they gradually came to resemble in a case of convergent evolution. Ichthyosaurs were particularly abundant in the Late Triassic a ...
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Callawayia
''Callawayia'' is an extinct genus of ichthyosaur. It contains the species ''Callawayia neoscapularis''.Sander, P.M., Chen X., Cheng L. and Wang X. (2011).Short-Snouted Toothless Ichthyosaur from China Suggests Late Triassic Diversification of Suction Feeding Ichthyosaurs. ''PLoS ONE 6(5)'': e19480. History of research In 1994, Christopher McGowan reviewed the taxonomy of '' Shastasaurus''. In this publication, he named a new species of ''Shastasaurus'', ''S. neoscapularis'', based on ROM (Royal Ontario Museum) 41993, a partial skeleton discovered on the shore of Williston Lake in Pink Mountain, British Columbia, Canada in 1987. The specimen was excavated from "Flipper Quarry" in 1988. The name of this species refers to the closer resemblance of its scapulae (shoulder blades) to Jurassic ichthyosaurs than those of other Triassic ichthyosaurs. In 2000, Michael Maisch published a study on the skull anatomy of ''Shastasaurus'', and also revised the genus again. He noted that ''S. ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Shastasaurus
''Shastasaurus'' ("Mount Shasta lizard") is a very large extinct genus of ichthyosaur from the middle and late Triassic, and is the largest known marine reptile.Hilton, Richard P., ''Dinosaurs and Other Mesozoic Animals of California'', University of California Press, Berkeley 2003 , at pages 90-91. Specimens have been found in the United States, Canada, and China. Description left, Size of '' Shonisaurus popularis'' (green) and ''S. sikanniensis'' (red), a possible species of ''Shastasaurus'', compared with a human ''Shastasaurus'' lived during the late Triassic period. The type species ''Shastasaurus pacificus'' is known from California. ''S. pacificus'' was a medium-sized ichthyosaur, measuring over in length and weighing . A second possible species of ''Shastasaurus'', ''S. sikanniensis'', is known from the Pardonet Formation British Columbia, dating to the middle Norian age (about 210 million years ago). If ''S.sikanniensis'' belongs to ''Shastasaurus'', it would be the la ...
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Pisiform Bone
The pisiform bone ( or ), also spelled pisiforme (from the Latin ''pisifomis'', pea-shaped), is a small knobbly, sesamoid bone that is found in the wrist. It forms the ulnar border of the carpal tunnel. Structure The pisiform is a sesamoid bone, with no covering membrane of periosteum. It is the last carpal bone to ossify. The pisiform bone is a small bone found in the proximal row of the wrist (carpus). It is situated where the ulna joins the wrist, within the tendon of the flexor carpi ulnaris muscle. It only has one side that acts as a joint, articulating with the triquetral bone. It is on a plane anterior to the other carpal bones and is spheroidal in form. The pisiform bone has four surfaces: # The ''dorsal surface'' is smooth and oval, and articulates with the triquetral: this facet approaches the superior, but not the inferior border of the bone. # The ''palmar surface'' is rounded and rough, and gives attachment to the transverse carpal ligament, the flexor carpi ulnaris ...
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Coracoid
A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is not homologous with the coracoid bone of most other vertebrates. In other tetrapods it joins the scapula to the front end of the sternum and has a notch on the dorsal surface which, along with a similar notch on the ventral surface of the scapula, forms the socket in which the proximal end of the humerus (upper arm bone) is located. The acrocoracoid process is an expansion adjacent to this contact surface, to which the shoulderward end of the biceps brachii muscle attaches in these animals. In birds (and generally theropods and related animals), the entire unit is rigid and called scapulocoracoid. This plays a major role in bird flight. In dinosaurs the main bones of the pectoral girdle were the sca ...
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Scapula
The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side of the body being roughly a mirror image of the other. The name derives from the Classical Latin word for trowel or small shovel, which it was thought to resemble. In compound terms, the prefix omo- is used for the shoulder blade in medical terminology. This prefix is derived from ὦμος (ōmos), the Ancient Greek word for shoulder, and is cognate with the Latin , which in Latin signifies either the shoulder or the upper arm bone. The scapula forms the back of the shoulder girdle. In humans, it is a flat bone, roughly triangular in shape, placed on a posterolateral aspect of the thoracic cage. Structure The scapula is a thick, flat bone lying on the thoracic wall that provides an attachment for three groups of muscles: intrin ...
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Pectoral Girdle
The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists of the clavicle, scapula, and coracoid. Some mammalian species (such as the dog and the horse) have only the scapula. The pectoral girdles are to the upper limbs as the pelvic girdle is to the lower limbs; the girdles are the parts of the appendicular skeleton that anchor the appendages to the axial skeleton. In humans, the only true anatomical joints between the shoulder girdle and the axial skeleton are the sternoclavicular joints on each side. No anatomical joint exists between each scapula and the rib cage; instead the muscular connection or physiological joint between the two permits great mobility of the shoulder girdle compared to the compact pelvic girdle; because the upper limb is not usually involved in weight bearing, its stabilit ...
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Toretocnemidae
The Toretocnemidae were a group of ichthyosaurs that lived in the Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ... period. References *Maisch, M.W. and Matzke, A.T., 2000, "The Ichthyosauria", ''Stuttgarter Beiträge zur Naturkunde Series B (Geologie und Paläontologie)'', 298: 1-159 Late Triassic ichthyosaurs Prehistoric reptile families {{triassic-reptile-stub ...
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Cymbospondylidae
Cymbospondylidae is an extinct family of Hueneosaurian Ichthyosaurs known from the Middle Triassic of Europe, North America, and Asia. Taxonomy Cymbospondylidae is a basal clade of ichthyosaurs. In 2000, Maischand and Matzke recovered cymbospondylids as members of Hueneosauria and more derived than mixosaurids, but Ji and colleagues found cymbospondylids to be outside of Hueneosauria in 2015. These two studies defined the group in two different ways, with the former defining it as the least inclusive clade containing both '' Cymbospondylus petrinus'' and '' Phantomosaurus neubigi'', whereas the latter defined it as the least inclusive clade containing both ''Cymbospondylus piscosus'' and ''Xinminosaurus catactes ''Xinminosaurus'' is an extinct genus of cymbospondylid ichthyosaur known from the Middle Triassic (mid-late Anisian stage) of Guizhou Province, China. Etymology The generic name is derived from ''Xinmin'', the district where the fossil was ...''. References ...
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Mixosauria
The Mixosauria were an early group of ichthyosaurs, living between 247.2 and 235 million years ago, during the Triassic period. Fossils of mixosaurs have been found all over the world: China, Timor, Indonesia, Italy, Germany, Spitsbergen, Switzerland, Svalbard, Canada, Alaska, and Nevada. Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... based on Motani (1999), Maisch and Matzke (2000), and Jiang, Schmitz, Hao & Sun (2006), with clade and generic names following Maisch (2010): References Triassic ichthyosaurs Triassic reptiles of Asia Triassic reptiles of Europe Triassic reptiles of North America Fossil taxa described in 1999 {{ichthyosaur-stub ...
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