Eodicynodontidae
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Eodicynodontidae
''Eodicynodon'' (''eo-,'' early or primitive, dicynodont) is an extinct genus of dicynodont therapsids, a highly diverse group of herbivorous synapsids that were widespread during the middle-late Permian and early Triassic. As its name suggests, ''Eodicynodon'' is the oldest and most primitive dicynodont yet identified, ranging from the middle to late Permian and possessing a mix of ancestral Anomodont/therapsid features and derived dicynodont synapomorphies. First described by paleontologist T. H. Barry in 1974, its only associated species, ''E. oosthuizeni'', is named after Roy Oosthuizeni, the South African farmer who discovered the type specimen (a partial skull without the mandible) on his Cape Province farm between 1964 and 1970. Description ''Eodicynodon'' was a medium-sized dicynodont, at about 450 mm long and 150 mm high. While it had many features common to all dicynodonts, such as canine tusks and jaw structures related to the "cheek pivot system" of ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Sphenacodontia
Sphenacodontia is a stem-based clade of derived synapsids. It was defined by Amson and Laurin (2011) as "the largest clade that includes ''Haptodus baylei'', ''Haptodus garnettensis'' and '' Sphenacodon ferox'', but not ''Edaphosaurus pogonias''". They first appear during the Late Pennsylvanian (Upper Carboniferous) epoch. From the end of the Carboniferous to the end of the Permian, most of them remained large, with only some secondarily becoming small in size. Basal Sphenacodontia constitute a transitional evolutionary series from early pelycosaurs to ancestral therapsids (which in turn were the ancestors of more advanced forms and finally the mammals). One might say that the sphenacodontians are proto-therapsids (even though there is almost a 30-million-years gap between the separation of the ancestors of therapsids from other sphenacodontians and the first appearance of therapsids in the fossil record). Characteristics The defining characteristics include a thickening of ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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Phthinosuchus
''Phthinosuchus'' is an extinct genus of therapsids from the Middle Permian of Russia. ''Phthinosuchus'' is the sole member of the family Phthinosuchidae. ''Phthinosuchus'' may have been one of the most primitive therapsids, meaning that its ancestors may have branched off early from the main therapsid line. Discovery ''Phthinosuchus'' was named in 1954 by Ivan Yefremov. It is currently only known from the back of the skull, as the front of the skull was lost after description. Two species, ''P. discors'' and ''P. horissiaki'', have been described. Description ''Phthinosuchus'' was 1.5 m (5 ft) long with a 20 cm skull, and looked much like the Sphenacodontids, such as ''Dimetrodon'' and ''Sphenacodon''. Its temporal fenestrae were larger than those of the Sphenacodontids. Its jaw was slender, unlike other predatory therapsids, but like the other early therapsids, it was probably sprawling and carnivorous. Classification ''Phthinosuchus'' is a member of the subor ...
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Eotitanosuchidae
Eotitanosuchidae is an extinct family of biarmosuchian therapsids. The Eotitanosuchidae were large predatory therapsids of the Wordian epoch. It was once considered to belong to a separate infraorder of therapsids called Eotitanosuchia. Characteristics The Eotitanosuchians seem to be more advanced than the Biarmosuchia in that the temporal opening behind the eye socket—although small—is still somewhat larger than the biarmosuchians; it is expanded in the upper rear ( posterodorsal) margin, allowing the area of attachment of the adductor ( jaw closing) muscles to be visible from the dorsal (top) view looking down. The eotitanosuchian bite was stronger and more efficient than the biarmosuchian bite. For this reason, some paleontologists see the eotitanosuchids as transitional between the biarmosuchians and higher therapsids. It is at least as likely that features of a larger temporal opening—and hence increased muscle mass and biting power—evolved simultaneously among a nu ...
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Biarmosuchia
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pr ...
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Tetraceratops
''Tetraceratops insignis'' ("four-horned face emblem") is an extinct synapsid from the Early Permian that was formerly considered the earliest known representative of Therapsida, a group that includes mammals and their close extinct relatives. It is known from a single skull, discovered in Texas in 1908. According to a 2020 study, it should be classified as a primitive non-therapsid sphenacodont rather than a genuine basal therapsid. Description ''Tetraceratops'' is known from a single skull discovered in Texas in the early 1900s. Contrary to its genus name, ''Tetraceratops'' actually has six horns, one pair being on the premaxilla bones, one pair on the prefrontal bones, and one pair on the angular processes of the mandible. When it was discovered and described in 1908, the skull was still embedded in a matrix, and only the premaxilla and prefrontal pairs were visible. In life, thus, it would have resembled a large lizard with four horns on its snout, and a pair of large s ...
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Sphenacodontidae
Sphenacodontidae (Greek: "wedge point tooth family") is an extinct family of small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian pelycosaurs. The most recent one, ''Dimetrodon angelensis'', is from the late Kungurian or early Roadian San Angelo Formation. However, given the notorious incompleteness of the fossil record, a recent study concluded that the Sphenacodontidae may have become extinct as recently as the early Capitanian. Primitive forms were generally small (60 cm to 1 meter), but during the later part of the early Permian these animals grew progressively larger (up to 3 meters or more), to become the top predators of their environments. Sphenacodontid fossils are so far known only from North America and Europe. Characteristics The skull is long, deep and narrow, an adaptation for strong jaw muscles. The front teeth are large and dagger-like, whereas the teeth in the sides and rear of the jaw are much smaller (hence the name of the well-kn ...
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Caseasauria
Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insectivorous or carnivorous Eothyrididae, and the large, herbivorous Caseidae. These two groups share a number of specialised features associated with the morphology of the snout and external naris. The ancestor of caseasaurs can be traced back to an insect eating or an omnivorous reptile-like synapsid from the Pennsylvanian time of the Carboniferous, possibly resembling ''Archaeothyris'', the earliest known synapsid. The caseasaurs were abundant during the later part of the Early Permian, but by the Middle Permian caseasaur diversity declined because the group was outcompeted by the more successful therapsids. The last caseasaurs became extinct at the end of the Guadelupian (Middle Permian). Description Among the most conspicuous charact ...
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Ecca Group
The Ecca Group is the second of the main subdivisions of the Karoo Supergroup of geological strata in southern Africa. It mainly follows conformably after the Dwyka Group in some sections, but in some localities overlying unconformably over much older basement rocks. It underlies the Beaufort Group in all known outcrops and exposures. Based on stratigraphic position, lithostratigraphic correlation, palynological analyses, and other means of geological dating, the Ecca Group ranges between Early to earliest Middle Permian (Asselian - Roadian) in age. Background During the time of the deposition of the Ecca Group, the depositional environment, with some exceptions, was predominantly marine. The Ecca sea was vast but shallow, reaching only around 500 m at its deepest in its west/northwestern and southern facies where the Tanqua and Laingsburg Depocenters are situated respectively. The marine environment ranged from deep pelagic, submarine fan systems in the lower deposits whi ...
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Beaufort Group
The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlies the Stormberg Group. Based on stratigraphic position, lithostratigraphic and biostratigraphic correlations, palynological analyses, and other means of geological dating, the Beaufort Group rocks are considered to range between Middle Permian (Wordian) to Early Triassic (Anisian) in age. Background During the period when sedimentation of the Beaufort Group rocks took place, the Ecca sea had retreated to the northeastern Karoo Basin. All sediment deposition at this time took place in a terrestrial, although in a predominantly fluvial or alluvial environment that was seasonally arid. This environment covered a vast area and deposition was influenced by a retroarc foreland basin. This foreland system was caused by crustal uplift (oro ...
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Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of the Mesozoic, Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic magmatic province, Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by an extinction event associated with widespread Anoxic event, oceanic anoxia, ocean acidification, and elevated temperatures likely caused by the eruption of the Karoo-Ferrar, Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and i ...
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