Ellesmeroceratida
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Ellesmeroceratida
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that are typically small, some even tiny, with close-spaced septa and relatively large ventral siphuncles. In some genera (e.g. ''Paleoceras''), the septa are uniformly spaced. Shells of ellesmerocerids are typically smooth and compressed and vary in form. They may be breviconic (short) or longiconic (elongate), straight (orthoconic) or curved (cyrtoconic). Cyrtoconic forms are usually endogastric, with longitudinally convex ventral margins. The apeces of straight forms typically have an endogastric curvature. Some may have grown to as much as 15 cm. Siphuncle segments are tubular or concave. Septal necks are short. Connecting rings which may appear layered are thick and typically wedge shaped with their maximum width at or near where t ...
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Ellesmeroceratidae
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic). Evolution and phylogeny The Ellesmeroceratidae have their derivation in the Plectronoceratidae, order Plectronocerida, in Tr ...
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Nautiloidea
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Ectenolites
''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' Ellesmeroceras'' but are smaller and proportionally narrower. Septa, as typical for ellesmerocerids, are close spaced with shallow lobes on either flank. The body chamber is proportionally long, the shell itself slightly compressed. The dorsal side at the beginning of the shell, opposite the aperture and body chamber, is strongly convex so to produce a sense of endogastric curvature with the apex and siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... aligned.Flower R.H.1964, The Nautiloid Order Ellesmeroceratida (Cephalopoda). Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, NM The siphuncle in ''Ectenolites'' lies along the ventral side; is tubular, composed ...
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Plectronocerida
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.Curt Teichert, 1988. Main Features of Cephalopod Evolution. The Mollusca Vol. 12 Paleontology and Neontology of Cephalopds; Academic Pres Inc. Occurrence Plectronoceratids are known from the Upper Cambrian (upper Franconian – middle Trempealeauan) of China and North America (Minnesota, Wisconsin). Two families are recognized (Flower, 1964),Flower, Rousseau H. 1964. The Nautiloid Order Ellesmeroceratida (Cephalopoda); Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, New Mexico the generally straight to endogastric Plectronoceratidae and the slightly exogastric Balkoceratidae. Diagnostic characters Members of the Plectronocerida are characterized as follows. Shells are generally small, some even tiny, laterally compressed, curved (cyrtochonic) or straight (orthoconic). Most cyrtoconic forms are endogastric, with the ventral side longitudinally concave, or t ...
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Protocycloceratidae
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician. Protocycloceratidae, named by Kobayashi (1935), are characterized by transversely ribbed, elongate shells that may be straight or slightly curved, as to be either endogastric or exogastric.Flower 1964, Furnish & Glenister 1964 Although not universally accepted, Kröger and Mutvie (2005) revised the Protocycloceratidae to include only those ellesmerocerids that have straight transverse sutures and siphuncles with concave segments and thick connecting rings. Protocycloceratidae (''sensu'' Flower, 1964) includes the follow genera, given with brief descriptions. *'' Protocycloceras''. Strongly annulate orthocones; cross section circular; annulations and sutures transverse and essentially straight; siphuncle tubular, ventral but not marginal, about 0.3 corresponding shell diameter. Lower Ordovician, cosmopolitan. *'' Catoraphi ...
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Gasconadian Stage
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Bathmoceratidae
''Bathmoceras'' is a primitive cephalopod genus from the Middle and Upper Ordovician. It is a member of the order Cyrtocerinida and is the only genus in the family Bathmoceratidae. Derivation At one time ''Bathmoceras'' was thought to have possibly given rise to ''Polydesmia'', once thought to have been the ancestral actinocerid. Since then revision of actinocerid phylogeny and of Ordovician stratigraphy in east Asia have cast doubts on this hypothesis. ''Bathmoceras'' is most likely derived from ''Eothinoceras'' through lengthening of the septal necks and of the inward projections of the connecting rings in a forward direction. ''Eothinoceras'' differs in that the septal necks are vestigial rather than being somewhat long as in ''Bathmoceras'' and that the inward projections of the connecting rings point straight in rather than projecting forward. Both are straight shelled or slightly exogastric. Eothinoceras is also apparently more slender. Furnish and Glenister also include ...
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Cyrtocerinidae
Cyrtocerinidae is a family of nautiloid cephalopods in the Order Cyrtocerinida Cyrtocerinida is an order of Ordovician nautiloid cephalopods. The order includes the families Cyrtocerinidae and Eothinoceratidae, as well as the genera '' Bathmoceras'' and '' Rummoceras''. Cyrtocerinids can be characterized by a broad si ..., previously considered ellesmerocerids. Members of the family have slightly endogastric (inwardly curved) breviconic shells with ventral siphuncles that have connecting rings thickened as lobes that project straight into the interior. Cyrtocerinidae was established by Flower 1964 for the genus '' Cyrtocerina'' Billings 1865, found at various locations across North America. The shells are generally small, laterally compressed, and dorsum more narrowly rounded than the venter. References Prehistoric nautiloid families Ordovician first appearances Ordovician extinctions Ellesmerocerida {{paleo-nautiloidea-stub ...
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Apocrinoceratidae
Apocrinoceratidae constitutes a family of Middle Ordovician nautiloid cephalopods characterized by straight or slightly curved, transversely ribbed shells having siphuncles composed of expanded segments, short recurved septal necks, and thick connecting rings. Derivation is from the Protocycloceratidae, a family of ellesmerocerids, which differ in having straight or concave siphuncle segments, but are otherwise similar in form. Apocrinoceratidae was established by Rousseau Flower (in Flower & Teichert, 1957) repeated in Flower (1964) for the genus '' Apocrinoceras'' (Teichert & Glenister, 1954). Four additional genera are included; '' Desioceras'' and '' Glenisteroceras'' (Flower and Teichert 1957, Flower 1964),'' Bakeroceras'' (Hook and Flower 1977), and '' Paldoceras'' (Kröger ''et al'' 2009). Apocrinoceratidae was removed from the Ellesmerocerida to the Discosorida Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the ...
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Bassleroceras
''Bassleroceras'' is an elongate upwardly curved, exogastric, genus with the venter on the under side more sharply rounded than the dorsum on the upper. The siphuncle is ventral, composed of thick-walled tubular segments in which connection rings thicken in towardly as in both the Ellesmerocerida and primitive Tarphycerida. ''Bassleroceras'' is the type genus of the Bassleroceratidae which Furnish and Glenister (1964) included in the Ellesmerocerida, but which Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ... (1967) placed in the Tarphycerida. ''Bassleroceras'' gave rise to the Tarphycerida (sensu Furnish and Glenister, 1964) by evolving genera with tighter and tighter curvatures until becoming gyroconic, a character of the Estonioceratidae, a family of early tarphycerids ...
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Oncocerida
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are thin and siphuncle segments are variably expanded (Flower, 1950). At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three (Sweet, 1964). Physical characteristics The shells of oncocerids are primarily somewhat compressed cyrtoconic brevicones. More advanced forms include gyrocones, serpenticones, torticones, and elongate orthocones and cyrtocones, reflective of the different families and genera (Flower, 1950; Sweet, 1964). The siphuncle in the Oncocerida is commonly located at or near the ventral margin. Connecting rings are mo ...
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Tarphycerida
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician (middle and upper Canad) to the Middle Devonian. Some, such as '' Aphetoceras'' and '' Estonioceras'', are loosely coiled and gyroconic; others, such as '' Campbelloceras'', '' Tarphyceras'', and '' Trocholites'', are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular,Furnish and Glenister 1964; Nautiloidea - Tarphycerida; In the ''Treatise on Invertebrate Paleontology'' Vol K; Teichert and Moore, (eds) GSA and U of Kansas Press 1964 as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably ''Bassleroceras'', around the end of the Gasconadian through forms like ''Aphetoceras''. Close coiling developed rather quickly, and both gyroconic and evolute forms are fou ...
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