Plectronocerida
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Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.Curt Teichert, 1988. Main Features of Cephalopod Evolution. The Mollusca Vol. 12 Paleontology and Neontology of Cephalopds; Academic Pres Inc.


Occurrence

Plectronoceratids are known from the Upper Cambrian (upper Franconian – middle
Trempealeauan The Trempealeauan is the upper or latest stage of the Upper or Late Cambrian in North America, spanning about 4 million years from about 492.5 to 488.3 m.y.a., equivalent to the Fengshanian of China. The name comes from the Trempealeau Formation, ...
) of China and North America (Minnesota, Wisconsin). Two families are recognized (Flower, 1964),Flower, Rousseau H. 1964. The Nautiloid Order Ellesmeroceratida (Cephalopoda); Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, New Mexico the generally straight to endogastric
Plectronoceratidae Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.Curt Teichert, 1988. Main Features of Cephalopod Evolution. The Mollusca Vol. 12 Paleontology and Neontology of Cephalopds; Academic Pres Inc. Oc ...
and the slightly exogastric Balkoceratidae.


Diagnostic characters

Members of the Plectronocerida are characterized as follows. Shells are generally small, some even tiny, laterally compressed, curved (cyrtochonic) or straight (orthoconic). Most cyrtoconic forms are endogastric, with the ventral side longitudinally concave, or the dorsal side more longitudinally convex. A few, the two known genera in Balkoceratidae are exogastrically curved, with the ventral side convex and dorsal side concave. Septa are close spaced, in some less than a millimeter. Siphuncles are ventral, and in most, proportionally large. Connecting rings are in general poorly calcified and may expand as siphonal bulbs into the chambers where not restricted by septal necks. As with all shelled cephalopods, plectronocerids had a tube called a siphuncle, which let them fill the chambers of their
phragmocone The phragmocone is the chambered portion of the shell of a cephalopod. It is divided by septa into camerae. In most nautiloids and ammonoids, the phragmocone is a long, straight, curved, or coiled structure, in which the camarae are linked by a ...
with gas instead of water, thus controlling their buoyancy. They were not, however, adapted for jet-powered swimming.


Phylogeny

The Plectronoceratida gave rise in the
Late Cambrian Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
(early and middle Trempealeauan) to the other three plectronoceratoid orders, the
Ellesmerocerida The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that ...
and
Protactinocerida Protactinocerida is a proposed order of Late Cambrian nautiloid Cephalopod, cephalopods. Their fossils have only been found in the Late Cambrian (late Jiangshanian to early Cambrian Stage 10, Stage 10) of North China Craton, North China, a diversi ...
and Yanhecerida. The Plectronoceratidae gave rise to the other ellesmerocerid families, including the Upper Cambrian exogastric Balkoceratidea, and with remote possibility to the
Discosorida Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through an ...
. The Balkoceratidae are unrelated to later exogastric forms that first appeared in the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...
.


Ecology

Plectronocerids were probably benthic animals that crawled along the bottom in search of food or safety, facing downwards, with the shell carried above. Nothing is known of their specific soft-part anatomy or to what extent tentacles, if any, had developed.


References


Further reading

* * {{Taxonbar, from=Q3906621 Prehistoric cephalopod orders Prehistoric nautiloids Cambrian animals of Asia Cambrian animals of North America Cambrian first appearances Furongian extinctions Paraphyletic groups