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Erythrovenator
''Erythrovenator'' () is a genus of basal theropod dinosaurs from the Late Triassic of Rio Grande so Sul, Brazil. The genus contains a single species, ''Erythrovenator jacuiensis''. Discovery The holotype and only known specimen of ''Erythrovenator'', CAPPA/UFSM 0157, is an isolated proximal portion of the left femur. This fossil was found in red mudstone of the Niemeyer Site, near Agudo in Rio Grande Sul. The site is tentatively considered to be early Norian (or possible late Carnian) in age and belongs the Candelária Sequence of the Santa Maria Supersequence. No other dinosauromorphs are known from this site, which is dominated by the traversodontid cynodont '' Siriusgnathus''. The site probably corresponds to the ''Riograndia'' Assemblage Zone based on the presence of ''Siriusgnathus''. The generic name, ''Erythrovenator'', is derived from the Greek word ερυθρός (''erythrós''), meaning "red" (in reference to the color of the holotype), and the Latin w ...
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Erythrovenator Jacuiensis
''Erythrovenator'' () is a genus of basal theropod dinosaurs from the Late Triassic of Rio Grande so Sul, Brazil. The genus contains a single species, ''Erythrovenator jacuiensis''. Discovery The holotype and only known specimen of ''Erythrovenator'', CAPPA/UFSM 0157, is an isolated proximal portion of the left femur. This fossil was found in red mudstone of the Niemeyer Site, near Agudo in Rio Grande Sul. The site is tentatively considered to be early Norian (or possible late Carnian) in age and belongs the Candelária Sequence of the Santa Maria Supersequence. No other dinosauromorphs are known from this site, which is dominated by the traversodontid cynodont '' Siriusgnathus''. The site probably corresponds to the ''Riograndia'' Assemblage Zone based on the presence of ''Siriusgnathus''. The generic name, ''Erythrovenator'', is derived from the Greek word ερυθρός (''erythrós''), meaning "red" (in reference to the color of the holotype), and the Latin word ...
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Candelária Formation, Paraná Basin
The Candelária Formation, in other literature also referred to as Candelária Sequence,Müller et al., 2017, p.543Martinelli et al., 2017, p.528 is a sedimentary formation of the Santa Maria Group (also called Santa Maria Supersequence) in the Paraná Basin in Rio Grande do Sul, southeastern Brazil.Candelária Formation
at .org
The formation dates to the of the

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Theropoda
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered ...
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Late Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and '' Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', the middle Muschelkalk (shell-bearing limestone) and the upper Keuper (coloured clay). The Late Triassic Ser ...
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Traversodontidae
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil. Description Traversodonts are members of Gomphodontia, a group of herbivorous cynognathian cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large canines. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of traverso ...
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Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, the fourth-largest country in the Americas, and the eighth-largest country in the world. It shares the bulk of the Southern Cone with Chile to the west, and is also bordered by Bolivia and Paraguay to the north, Brazil to the northeast, Uruguay and the South Atlantic Ocean to the east, and the Drake Passage to the south. Argentina is a federal state subdivided into twenty-three provinces, and one autonomous city, which is the federal capital and largest city of the nation, Buenos Aires. The provinces and the capital have their own constitutions, but exist under a federal system. Argentina claims sovereignty over the Falkland Islands, South Georgia and the South Sandwich Islands, and a part of Antarctica. The earliest recorded human presen ...
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Eodromaeus
''Eodromaeus'' (meaning "dawn runner") is an extinct genus of probable basal theropod dinosaurs from the Late Triassic of Argentina. Like many other of the earliest-known dinosaurs, it hails from the Carnian-age (~230 Ma) Ischigualasto Formation, within the Ischigualasto-Villa Unión Basin of northwestern Argentina. Upon its discovery, it was argued to be one of the oldest true theropods, supplanting its contemporary '' Eoraptor'', which was reinterpreted as a basal sauropodomorph.Martínez et al., 2011 Discovery Fossils from ''Eodromaeus'' were first discovered in 1996 by Argentinean paleontologist Ricardo N. Martinez and Earthwatch volunteer Jim Murphy, and it was first believed that the fossils were a new species of '' Eoraptor''. However, as the researchers started to take a closer look at the fossils, they found that it had many skeletal features which were absent in ''Eoraptor'', and they understood that it came from a new genus. ''Eodromaeus'' is known from six ...
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Silesauridae
Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids comprised an early diverging clade or a paraphyletic grade within ornithischian dinosaurs. Silesaurids have a consistent general body plan, with a fairly long neck and legs and possibly quadrupedal habits, but most silesaurids are heavily fragmentary nonetheless. Furthermore, they occupied a variety of ecological niches, with early silesaurids (such as ''Lewisuchus'') being carnivorous and later taxa (such as ''Kwanasaurus'') having adaptations for specialized herbivory. As indicated by the contents of referred coprolites, ''Silesaurus'' may have been insectivorous, feeding selectively on small beetles and other arthropods. Classification Silesauridae is typically considered the sister group to Dinosauria. The group was named in 2010 by Max ...
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Neotheropoda
Neotheropoda (meaning "new theropods") is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the end of the Early Jurassic ( Pliensbachian) period except for Averostra. Classification Neotheropoda was named by Robert T. Bakker in 1986 as a group including the relatively derived theropod subgroups Ceratosauria and Tetanurae, and excluding coelophysoids. However, most later researchers have used it to denote a broader group. Neotheropoda was first defined as a clade by Paul Sereno in 1998 as '' Coelophysis'' plus modern birds, which includes almost all theropods except the most primitive species.Sereno, 1998. A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 210, 41-83. Dilophosauridae was formerly considered a sma ...
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Buriolestes
''Buriolestes'' is a genus of early sauropodomorph dinosaurs from the Late Triassic Santa Maria Formation of the Paraná Basin in southern Brazil. It contains a single species, ''B. schultzi'', named in 2016. The type specimen was found alongside a specimen of the lagerpetid dinosauromorph '' Ixalerpeton''. Discovery and naming The holotype specimen, ULBRA-PVT280, was discovered in 2009 in the Buriol ravine in São João do Polêsine, Brazil. These rocks are part of the Santa Maria Formation, which dates to the Carnian epoch. The specimen consists of a single skeleton preserving parts of the skull, vertebrae, left forelimb, and left hindlimb. Another set of smaller bones is also present, which may belong to a juvenile or a different taxon altogether. Two individuals of '' Ixalerpeton'' were also preserved close by. Additional remains were discovered in 2015 and published in 2018. These include a mostly complete skeleton, CAPPA/UFSM 0035, which preserves a complete skull and ...
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Jacuí River
The Jacuí River () is a river in Rio Grande do Sul state of southern Brazil. The Jacuí empties into the Guaíba River, an estuarine arm of the Lagoa dos Patos, a large coastal lagoon connected to the Atlantic Ocean. Sources The Jacuí River, known as 'Rio Jacuí' in Portuguese, has its origins in the highlands east of Passo Fundo. From there it flows south and then east for nearly . The Taquari, Caí, Sinos, and Gravataí rivers merge into the Jacuí near its mouth. At Porto Alegre, near the Atlantic coast, the Jacuí transforms into a shallow estuary, known as the Guaíba River, and flows into the Patos Lagoon. Boats can travel up the river as far as Cachoeira do Sul. Regional effects The Dona Francisca Hydroelectric Dam impounds the upper Jacuí River, and is divided between the municipalities of Agudo and Nova Palma Nova Palma is a municipality in the state of Rio Grande do Sul, Brazil. The municipality contains part of the hydroelectric Dona Francisca Dam on the u ...
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