Epioblasma Torulosa
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Epioblasma Torulosa
''Epioblasma torulosa'', commonly called the tubercled blossom, is a species of freshwater mussel, a mollusk in the family Unionidae. It is native to eastern North America, where it is considered endangered in both Canada and the United States. Natural history This species is known from the Cumberland, Ohio, St. Lawrence, and Tennessee River drainages. It is found in shallow riffle zones over sand and gravel in large to medium sized rivers. Due to habitat destruction, siltation, and other factors, two out of the three subspecies are now extinct. The surviving subspecies is considered highly imperiled, with only three known locations showing evidence of reproduction. Subspecies This species exhibits variable shell characteristics across its geographic range. It is unclear whether this is best understood as ecophenotypic variation, various subspecies, or a complex of closely related but genetically isolated species. Due to the extinction of three of the four subspecies often inclu ...
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Constantine Samuel Rafinesque
Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimately settling in Ohio in 1815, where he made notable contributions to botany, zoology, and the study of prehistoric earthworks in North America. He also contributed to the study of ancient Mesoamerican linguistics, in addition to work he had already completed in Europe. Rafinesque was an eccentric and erratic genius. He was an autodidact, who excelled in various fields of knowledge, as a zoologist, botanist, writer and polyglot. He wrote prolifically on such diverse topics as anthropology, biology, geology, and linguistics, but was honored in none of these fields during his lifetime. Indeed, he was an outcast in the American scientific community whose submissions were rejected automatically by leading journals. Among his theories were th ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Freshwater Mussel
Freshwater bivalves are one kind of freshwater mollusc, along with freshwater snails. They are bivalves that live in fresh water as opposed to salt water, which is the main habitat type for bivalves. The majority of species of bivalve molluscs live in the sea, but in addition, a number of different families live in fresh water (and in some cases, also in brackish water). These families belong to two different evolutionary lineages (freshwater mussels and freshwater clams), and the two groups are not closely related. Freshwater bivalves have a simple morphology that varies among taxa, and are distributed around most regions of the world. Species in the two groups vary greatly in size. Some pea clams (''Pisidium'' species) have an adult size of only 3 mm. In contrast, one of the largest species of freshwater bivalves is the swan mussel, in the family Unionidae; it can grow to a length of 20 cm, and usually lives in lakes or slow rivers. Freshwater pearl mussels are eco ...
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Mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gas ...
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Unionidae
The Unionidae are a family of freshwater mussels, the largest in the order Unionida, the bivalve molluscs sometimes known as river mussels, or simply as unionids. The range of distribution for this family is world-wide. It is at its most diverse in North America, with about 297 recognised taxa, but China and Southeast Asia also support very diverse faunas. Freshwater mussels occupy a wide range of habitats, but most often occupy lotic waters, i.e. flowing water such as rivers, streams and creeks. Origin and early diversification The recent phylogenetic study reveals that the Unionidae most likely originated in Southeast and East Asia in the Jurassic, with the earliest expansions into North America and Africa (since the mid-Cretaceous) followed by the colonization of Europe and India (since the Paleocene). Life history Unionidae burrow into the substrate, with their posterior margins exposed. They pump water through the incurrent aperture, obtaining oxygen and food. They remove ...
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Cumberland River
The Cumberland River is a major waterway of the Southern United States. The U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed June 8, 2011 river drains almost of southern Kentucky and north-central Tennessee. The river flows generally west from a source in the Appalachian Mountains to its confluence with the Ohio River near Paducah, Kentucky, and the mouth of the Tennessee River. Major tributaries include the Obey, Caney Fork, Stones, and Red rivers. Although the Cumberland River basin is predominantly rural, there are also some large cities on the river, including Nashville and Clarksville, both in Tennessee. Also, the river system has been extensively altered for flood control. Major dams impound areas of both the main stem and many of its important tributaries. Geography Its headwaters are three separate forks that begin in Kentucky and converge in Baxter, KY, located in Harlan County. Martin's Fork starts n ...
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Ohio River
The Ohio River is a long river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing southwesterly from western Pennsylvania to its mouth on the Mississippi River at the southern tip of Illinois. It is the third largest river by discharge volume in the United States and the largest tributary by volume of the north-south flowing Mississippi River that divides the eastern from western United States. It is also the 6th oldest river on the North American continent. The river flows through or along the border of six states, and its drainage basin includes parts of 14 states. Through its largest tributary, the Tennessee River, the basin includes several states of the southeastern U.S. It is the source of drinking water for five million people. The lower Ohio River just below Louisville is obstructed by rapids known as the Falls of the Ohio where the elevation falls in restricting larger commercial navigation, although in the 18th ...
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Tennessee River
The Tennessee River is the largest tributary of the Ohio River. It is approximately long and is located in the southeastern United States in the Tennessee Valley. The river was once popularly known as the Cherokee River, among other names, as the Cherokee people had their homelands along its banks, especially in what are now East Tennessee and northern Alabama. Additionally, its tributary, the Little Tennessee River, flows into it from Western North Carolina and northeastern Georgia, where the river also was bordered by numerous Cherokee towns. Its current name is derived from the Cherokee town, ''Tanasi'', which was located on the Tennessee side of the Appalachian Mountains. Course The Tennessee River is formed at the confluence of the Holston and French Broad rivers in present-day Knoxville, Tennessee. From Knoxville, it flows southwest through East Tennessee into Chattanooga before crossing into Alabama. It travels through the Huntsville and Decatur area before rea ...
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Habitat Destruction
Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby reducing biodiversity and species abundance. Habitat destruction is the leading cause of biodiversity loss. Fragmentation and loss of habitat have become one of the most important topics of research in ecology as they are major threats to the survival of endangered species. Activities such as harvesting natural resources, industrial production and urbanization are human contributions to habitat destruction. Pressure from agriculture is the principal human cause. Some others include mining, logging, trawling, and urban sprawl. Habitat destruction is currently considered the primary cause of species extinction worldwide. Environmental factors can contribute to habitat destruction more indirectly. Geological processes, climate change, introdu ...
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Siltation
Siltation, is water pollution caused by particulate terrestrial clastic material, with a particle size dominated by silt or clay. It refers both to the increased concentration of suspended sediments and to the increased accumulation (temporary or permanent) of fine sediments on bottoms where they are undesirable. Siltation is most often caused by soil erosion or sediment spill. It is sometimes referred to by the ambiguous term "sediment pollution", which can also refer to a chemical contamination of sediments accumulated on the bottom, or to pollutants bound to sediment particles. Although "siltation" is not perfectly stringent, since it also includes particle sizes other than silt, it is preferred for its lack of ambiguity. Causes The origin of the increased sediment transport into an area may be erosion on land or activities in the water. In rural areas, the erosion source is typically soil degradation by intensive or inadequate agricultural practices, leading to soil eros ...
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Ecophenotypic Variation
Ecophenotypic variation ("ecophenotype") refers to phenotypical variation as a function of life station. In wide-ranging species, the contributions of heredity and environment are not always certain, but their interplay can sometimes be determined by experiment. Plants Plants display the most obvious examples of ecophenotypic variation. One example are trees growing in the woods developing long straight trunks, with branching crowns high in the canopy, while the same species growing alone in the open develops a spreading form, branching much lower to the ground. Genotypes often have much flexibility in the modification and expression of phenotypes; in many organisms these phenotypes are very different under varying environmental conditions. The plant ''Hieracium umbellatum'' is found growing in two different habitats in Sweden. One habitat is rocky sea-side cliffs, where the plants are bushy with broad leaves and expanded inflorescences; the other is among sand dunes where the pla ...
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Epioblasma Cincinnatiensis
''Epioblasma'' is a genus of freshwater mussels, aquatic bivalve mollusks in the family Unionidae, the river mussels. Most of the species in this genus have been lost in modern times, and the entire genus is threatened with the possibility of extinction. Reproduction All Unionidae are known to use the gills, fins, or skin of a host fish for nutrients during the larval glochidia stage. It was discovered in 2004 that female ''Epioblasma'' in the subgenus ''Torulosa'' transfer their parasitic larvae to the host fish by snapping onto the head of the fish and pumping the larvae into the host fish's gills. While using bait to lure host fish towards the larvae is common in the family Unionidae, this was the first time that "fish snapping" behavior had been observed. Examination of other species within the genus ''Epioblasma'' may further reveal unusual reproductive mechanisms. Taxonomy of the genus ''Epioblasma'' Note: Taxa with a " †" symbol are extinct due to human activity *Subgen ...
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