Epichloë
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Epichloë
''Epichloë'' is a genus of ascomycete fungi forming an endophytic symbiosis with grasses. Grass choke disease is a symptom in grasses induced by some ''Epichloë'' species, which form spore-bearing mats ( stromata) on tillers and suppress the development of their host plant's inflorescence. For most of their life cycle however, ''Epichloë'' grow in the intercellular space of stems, leaves, inflorescences, and seeds of the grass plant without incurring symptoms of disease. In fact, they provide several benefits to their host, including the production of different herbivore-deterring alkaloids, increased stress resistance, and growth promotion. Within the family Clavicipitaceae, ''Epichloë'' is embedded in a group of endophytic and plant pathogenic fungi, whose common ancestor probably derived from an animal pathogen. The genus includes both species with a sexually reproducing (teleomorphic) stage and asexual, anamorphic species. The latter were previously placed in the ...
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Epichloë Typhina
''Epichloë typhina'' is a haploid sexual species in the fungal genus ''Epichloë''. It was Species description, originally described as a ''Sphaeria'' species. Today, however, it is classified in ''Epichloë''. A systemic grass symbiont first species description, described as ''Sphaeria typhina'' by Christian Hendrik Persoon in 1798, and transferred to genus ''Epichloë'' in 1865, ''Epichloë typhina'' forms an only partially resolved species complex within the ''Epichloë'' genus. ''Epichloë typhina'' is found in Europe, but has been introduced widely in North America and elsewhere. It lives in association with a large number of grass species from multiple genera, including ''Anthoxanthum odoratum'', ''Brachypodium'' ''phoenicoides'', ''Brachypodium pinnatum'', ''Dactylis glomerata'', ''Lolium perenne'', ''Milium effusum'', Timothy-grass, ''Phleum pratense'', ''Poa trivialis'', ''Poa'' ''silvicola'' and ''Puccinellia distans''. ''Epichloë typhina'' appears to be seed-tran ...
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Fungus
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''true f ...
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Neotyphodium
''Neotyphodium'' is a genus of endophytic fungi symbiotic with grasses. It used to contain a number of asexually reproducing species that colonize the leaves of cool-season grasses, but most of them, including the type species '' N. coenophialum'', were merged into the genus ''Epichloë'' in 2014. Two species of unclear position were excluded from this treatment: *'' Neotyphodium chilense'' from Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ... should be treated as ''Acremonium chilense'' since the previous transfer to ''Neotyphodium'' is untested. *''Neotyphodium starrii'' is closely related to ''N. coenophialum'', but the taxonomic status within ''Epichloë'' (whether it is a distinct species or not) is unclear ('' nomen dubium''). References Hypocreales genera C ...
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Endophyte
An endophyte is an endosymbiont, often a bacterium or fungus, that lives within a plant for at least part of its life cycle without causing apparent disease. Endophytes are ubiquitous and have been found in all species of plants studied to date; however, most of the endophyte/plant relationships are not well understood. Some endophytes may enhance host growth, nutrient acquisition and improve the plant's ability to tolerate abiotic stresses, such as drought and decrease biotic stresses by enhancing plant resistance to insects, pathogens and herbivores. Although endophytic bacteria and fungi are frequently studied, endophytic archaea are increasingly being considered for their role in plant growth promotion as part of the core microbiome of a plant. History Endophytes were first described by the German botanist Johann Heinrich Friedrich Link in 1809. They were thought to be plant parasitic fungi and they were later termed as "microzymas" by the French scientist Béchamp. There wa ...
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Clavicipitaceae
The Clavicipitaceae are a family of fungi within the order Hypocreales. A 2008 estimate placed 43 genera in the family, but recent work has increased this number to 97. Phylogeny Molecular phylogenetic analysis of multigene DNA sequence data indicates the taxon Clavicipitaceae is paraphyletic, and consists of three well-defined clades, at least one of which is shared with members of another fungal family (Hypocreaceae). The evolution within the Clavicipitaceae is marked by interkingdom host jumping, and the range of this large and heterogeneous fungal group spans mutualistic plant symbionts, as well as parasites of plants, insects, and other fungi. Significance Many of its members produce alkaloids toxic to animals and humans. One of its most infamous species is ''Claviceps purpurea'', which has historical significance as the cause of St. Anthony's fire, also known as ergotism. Ergotism is caused by ergot alkaloids, such as ergotamine and ergocristine, which are chemical d ...
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Ryegrass
''Lolium'' is a genus of tufted grasses in the bluegrass subfamily (Pooideae). It is often called ryegrass, but this term is sometimes used to refer to grasses in other genera. They are characterized by bunch-like growth habits. ''Lolium'' is native to Europe, Asia and northern Africa, as well as being cultivated and naturalized in Australia, the Americas, and various oceanic islands. Ryegrasses are naturally diploid, with 2n=14, and are closely related to the fescues (''Festuca''). Ryegrass should not be confused with rye, which is a grain crop. Species Species of ''Lolium'' include: * ''Lolium arundinaceum'' (Schreb.) Darbysh. - Eurasia + North Africa from Portugal + Canary Islands to Himalayas + Xinjiang; naturalized in East Asia, Australia, North + South America, various islands * ''Lolium canariense'' Steud. - Canary Islands ryegrass - Canary Islands, Cape Verde * ''Lolium giganteum '' Lam. - Eurasia from Ireland to China; Bioko * ''Lolium hybridum'' Hausskn. - Assam, B ...
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Poa Pratensis
''Poa pratensis'', commonly known as Kentucky bluegrass (or blue grass), smooth meadow-grass, or common meadow-grass, is a perennial species of grass native to practically all of Europe, North Asia and the mountains of Algeria and Morocco. Although the species is spread over all of the cool, humid parts of the United States, it is not native to North America. The Spanish Empire brought the seeds of Kentucky bluegrass to the New World in mixtures with other grasses. In its native range, ''Poa pratensis'' forms a valuable pasture plant, characteristic of well-drained, fertile soil. It is also used for making lawns in parks and gardens and has established itself as a common invasive weed across cool moist climates like the Pacific Northwest and the Northeastern United States. When found on native grasslands in Canada, for example, it is considered an unwelcome exotic plant, and is indicative of a disturbed and degraded landscape. Taxonomy ''Poa pratensis'' was one of the many speci ...
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Food Web
A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one of two categories called trophic levels: 1) the autotrophs, and 2) the heterotrophs. To maintain their bodies, grow, develop, and to reproduce, autotrophs produce organic matter from inorganic substances, including both minerals and gases such as carbon dioxide. These chemical reactions require energy, which mainly comes from the Sun and largely by photosynthesis, although a very small amount comes from bioelectrogenesis in wetlands, and mineral electron donors in hydrothermal vents and hot springs. These trophic levels are not binary, but form a gradient that includes complete autotrophs, which obtain their sole source of carbon from the atmosphere, mixotrophs (such as carnivorous plants), which are autotrophic organisms that partially ...
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Plant Communities
A plant community is a collection or association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The components of each plant community are influenced by soil type, topography, climate and human disturbance. In many cases there are several soil types present within a given plant community. This is because the soil type within an area is influenced by two factors, the rate at which water infiltrates or exits (via evapotranspiration) the soil, as well as the rate at which organic matter (any carbon-based compound within the environment, such as decaying plant matter) enters or decays from the soil. Plant communities are studied substantially by ecologists, due to providing information on the effects of dispersal, tolerance to environmental conditions, and response to disturbance of a variety of plant species, information valuable to the comprehension of various plant ...
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Hybrid Speciation
Hybrid speciation is a form of speciation where hybridization between two different species leads to a new species, reproductively isolated from the parent species. Previously, reproductive isolation between two species and their parents was thought to be particularly difficult to achieve, and thus hybrid species were thought to be very rare. With DNA analysis becoming more accessible in the 1990s, hybrid speciation has been shown to be a somewhat common phenomenon, particularly in plants. In botanical nomenclature, a hybrid species is also called a nothospecies. Hybrid species are by their nature polyphyletic. Ecology A hybrid may occasionally be better fitted to the local environment than the parental lineage, and as such, natural selection may favor these individuals. If reproductive isolation is subsequently achieved, a separate species may arise. Reproductive isolation may be genetic, ecological, behavioral, spatial, or a combination of these. If reproductive isolation ...
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Tall Fescue
''Festuca arundinacea'' (syn., ''Schedonorus arundinaceus'' and ''Lolium arundinaceum'') is a species of grass commonly known as tall fescue. It is a cool-season perennial C3 species of bunchgrass native to Europe. It is an important forage grass throughout Europe, and many cultivars have been used in agriculture. It is also an ornamental grass in gardens, and a phytoremediation plant. The predominant cultivar found in British pastures is S170, an endophyte-free variety. In its native European environment, tall fescue is found in damp grasslands, river banks, and in coastal seashore locations. Its distribution is a factor of climatic, edaphic, or other environmental attributes. In New Zealand, where it is introduced, the species is particularly prolific in salt marshes, where it is often a major part of the plant biota. History Festuca arundinacea was originally developed in Kew Gardens in the United Kingdom.Cougnon et al. (2013). Performance and quality of tall fescue (Festuc ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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