Eophis
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Eophis
''Eophis'' is an early genus of stem-snake (clade Ophidia) containing one species, ''Eophis underwoodi'', from the Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ... (Bathonian) from the Forest Marble Formation of United Kingdom. It is known from a fragmentary dentary from Kirtlington Quarry. It was previously interpreted as an anguimorph lizard. The placement of ''Eophis'' as a stem snake was re-affirmed by subsequent analyses. Phylogeny Cladogram based in the phylogenetic analysis by Caldwell ''et al.'' (2015): See also References Prehistoric reptile genera Middle Jurassic reptiles of Europe Fossil taxa described in 2015 {{paleo-reptile-stub ...
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Ophidia
__FORCETOC__ Ophidia (also known as Pan-Serpentes) is a group of squamate reptiles including modern snakes and reptiles more closely related to snakes than to other living groups of lizards. Ophidia was defined as the "most recent common ancestor of ''Pachyrhachis'' and Serpentes (modern snakes), and all its descendants" by Lee and Caldwell (1998: 1551). The latter author has used Ophidia in a manner inconsistent with this definition, using it to incorporate other more basal stem-snakes, such as the Late Cretaceous ''Najash rioegrina'' or the Jurassic ''Diablophis'' and ''Portugalophis''. The clade name Ophidia derives from the Ancient Greek word (), meaning "small snake". Evolution Modern snakes are thought to have evolved from either burrowing or aquatic lizards during the mid-Cretaceous period, and the earliest known fossils date to around 112 Ma ago. However, the relationship between modern snake and more primitive snake ancestors, many of which retained hind limbs, is l ...
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Parviraptor
''Parviraptor'' is a genus of stem-snake (clade Ophidia) containing one species, ''Parviraptor estesi'', from the Late Jurassic (Tithonian) or Early Cretaceous (Berriasian) Purbeck Limestone Formation of Dorset, England. A second species, ''Parviraptor gilmorei'', was described from the Late Jurassic Morrison Formation of Western North America;Foster, J. (2007). "Table 2.1: Fossil Vertebrates of the Morrison Formation." ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World''. Indiana University Press. pp. 58-59. it was present in stratigraphic zone 4.Foster, J. (2007). "''Enneabatrachus hechti''" ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World''. Indiana University Press. p. 137. However, the second species was subsequently transferred to a separate genus '' Diablophis''. An indeterminate species is known from the Bathonian aged Kirtlington Mammal Bed ( Kilmaluag Formation, Scotland). Description The shape, length, and body form o ...
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Diablophis
''Diablophis'' is a genus of Late Jurassic stem-snake from the Morrison Formation of North America. The type and only species, ''D. gilmorei'' was once thought to be a species of ''Parviraptor ''Parviraptor'' is a genus of stem-snake (clade Ophidia) containing one species, ''Parviraptor estesi'', from the Late Jurassic (Tithonian) or Early Cretaceous (Berriasian) Purbeck Limestone Formation of Dorset, England. A second species, ''Pa ...'' but is now classified as its own genus. The animal is known from multiple specimens, the holotype being LACM 4684/140572, which consists of a broken right mandible, broken right maxilla and broken axis vertebrae. A number of other specimens have also been attributed to ''Diablophis'', including LACM 4684/140572 and LACM 5572/120732, the specimens previously attributed to ''Parviraptor'' and LACM 4684/120472. Phylogeny ''Diablophis'' has been recovered as a basal snake, though this placement is not undisputed. The cladogram from figure 4 ...
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Forest Marble Formation
The Forest Marble is a geological formation in England. Part of the Great Oolite Group, it dates to the late Bathonian stage of the Middle Jurassic.Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 538–541. . Lithology The primary lithology of the formation typically consists of greenish grey variably calcareous silicate mudstone, with lenticular cross bedded limestone units deposited in a marine setting. Dinosaurian fauna Ornithischians Saurischians Microvertebrate fauna Despite the formation being nearly entirely marine, at several localities abundant remains of terrestrial microvertebrates are found, the primary locality being the Kirtlington Mammal Bed (designated 3p) in Kirtlington Quarry near Kirtlington, Oxfordshire. Another important locality is Watton Cliff near Eype in Dorset. Amp ...
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Prehistoric Reptile Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Serpentes
Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have evolved elongate bodies without limbs or with greatly reduced limbs about twenty-five times independently via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaenia, Dibamida ...
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Coniophis
''Coniophis'' is an extinct genus of snakes from the late Cretaceous period. The type species, '' Coniophis precedes'', was about 7 cm long and had snake-like teeth and body form, with a skull and a largely lizard-like bone structure. It probably ate small vertebrates. The fossil remains of ''Coniophis'' were first discovered at the end of the 19th century in the Lance Formation of the US state of Wyoming, and were described in 1892 by Othniel Charles Marsh. For the genus ''Coniophis'', a number of other species have been described. Their affiliation is, however, poorly secured, mostly based on vertebrae descriptions from only a few fossils. Fossil distribution Fossils of ''Coniophis'' have been found in: ;Cretaceous * Milk River and Frenchman Formations, Canada (Alberta, Saskatchewan) * Intertrappean Beds, India * Wadi Milk Formation The Wadi Milk Formation is a geological formation in Sudan whose strata date back to the Late Cretaceous. Originally, the formati ...
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Najash
''Najash'' is an extinct genus of basal snake from the Late Cretaceous Candeleros Formation of Patagonia. Like a number of other Cretaceous and living snakes it retained hindlimbs, but ''Najash'' is unusual in having well-developed legs that extend outside the rib cage, and a pelvis connected to the spine. Discovery and Description Fossils of ''Najash'' were found in the terrestrial Candeleros Formation, in Rio Negro Province, Argentina, and date to roughly 90 million years ago. The skull and spine of ''Najash'' show primitive features that resemble other Cretaceous snakes, such as ''Dinilysia patagonica'' and Madtsoiidae. Also, several characteristics of the neck and tail of ''Najash'' and '' Dinilysia patagonica'' show how the body plan of snakes evolved from a lizard-like ancestor. ''Najash'' had not lost its sacrum, the pelvic bone composed of several fused vertebrae, nor its pelvic girdle, which are absent in modern snakes, and in all other known fossil snakes as wel ...
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Madtsoiidae
Madtsoiidae is an extinct family of mostly Gondwanan snakes with a fossil record extending from early Cenomanian (Upper Cretaceous) to late Pleistocene strata located in South America, Africa, India, Australia and Southern Europe. Madtsoiidae include very primitive snakes, which like extant boas and pythons would likely dispatch their prey by constriction. Genera include '' Madtsoia'', one of the longest snakes known, at an estimated , and the Australian '' Wonambi'' and '' Yurlunggur''. As a grouping of basal forms the composition and even the validity of Madtsoiidae is in a state of flux as new pertinent finds are described, with more recent evidence suggesting that it is paraphyletic as previously defined. Although madtsoiids persisted on Australia until the Pleistocene, they largely went extinct elsewhere during the Eocene. However, some species persisted in South America and India through the Oligocene. Description Madtsoiidae was first classified as a subfamily ...
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Bathonian
In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Stratigraphic definitions The Bathonian Stage takes its name from Bath, a spa town in England built on Jurassic limestone (the Latinized form of the town name is ''Bathonium''). The name was introduced in scientific literature by Belgian geologist d'Omalius d'Halloy in 1843. The original type locality was located near Bath. The French palaeontologist Alcide d'Orbigny was in 1852 the first to define the exact length of the stage. The base of the Bathonian is at the first appearance of ammonite species '' Parkinsonia (Gonolkites) convergens'' in the stratigraphic column. The global reference profile for the base of the Bathonian (a GSSP) was ratified as Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France in 2009. The top of the Bat ...
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Dinilysia
''Dinilysia'' (meaning "terrible ilysia") is an extinct genus of snake from the Late Cretaceous (Coniacian) of South America. ''Dinilysia'' was a relatively large ambush predator, measuring approximately long. The skull morphology of ''Dinilysia'' is similar to boids, suggesting that it was able to consume large prey. Living in a desert-like environment, ''Dinilysia'' is likely a terrestrial or a semi-fossorial A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric eviden ... animal. Physiology and Lineage The ''Dinilysia patagonica'' is a stem snake that is very closely related to the original ancestor of the clade of crown snakes. Once the fossil of the snake was discovered, an x-ray computed tomography was used to build a digitized endocast of its inner ear. The results di ...
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