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Endodermal Cells
The endodermis is the central, innermost layer of cortex in land plants. It is a cylinder of compact living cells, the radial walls of which are impregnated with hydrophobic substances (Casparian strip) to restrict apoplastic flow of water to the inside. The endodermis is the boundary between the cortex and the stele. In many seedless plants (like ferns and mosses), the endodermis is a distinctly visible layer of cells immediately outside the vascular cylinder (stele) in roots and shoots. In most seed plants, especially woody types, an endodermis is absent from the stems but is present in roots. The endodermis helps regulate the movement of water, ions and hormones into and out of the vascular system. It may also store starch, be involved in perception of gravity and protect the plant against toxins moving into the vascular system. Structure The endodermis is developmentally the innermost portion of the cortex. It may consist of a single layer of barrel-shaped cells without any ...
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Ranunculus Root Cross Section
''Ranunculus'' is a large genus of about almost 1700 to more than 1800 species of flowering plants in the family Ranunculaceae. Members of the genus are known as buttercups, spearworts and water crowfoots. The genus is distributed in Europe, North America and South America. The familiar and widespread buttercup of gardens throughout Northern Europe (and introduced elsewhere) is the creeping buttercup ''Ranunculus repens'', which has extremely tough and tenacious roots. Two other species are also widespread, the bulbous buttercup ''Ranunculus bulbosus'' and the much taller meadow buttercup ''Ranunculus acris''. In ornamental gardens, all three are often regarded as weeds. Buttercups usually flower in the spring, but flowers may be found throughout the summer, especially where the plants are growing as opportunistic colonizers, as in the case of garden weeds. The water crowfoots (''Ranunculus'' subgenus ''Batrachium''), which grow in still or running water, are sometimes trea ...
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Cortex (botany)
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Plants Stems and branches In the three dimensional structure of herbaceous stems, the epidermis, cortex and vascular cambium form concentric cylinders around the inner cylindrical core of pith. Some of the outer cortical cells may contain chloroplasts, giving them a green color. They can therefore produce simple carbohydrates through photosynthesis. In woody plants, the cortex is located between the periderm (bark) and the vascular tissue (phloem, in particular). It is responsible for the transportation of materials into the central cylinder of the root through diffusion and may als ...
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Land Plants
The Embryophyta (), or land plants, are the most familiar group of green plants that comprise vegetation on Earth. Embryophytes () have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of green algae as sister of the Zygnematophyceae. The Embryophyta consist of the bryophytes plus the polysporangiophytes. Living embryophytes therefore include hornworts, liverworts, mosses, lycophytes, ferns, gymnosperms and flowering plants. The land plants have diplobiontic life cycles and it is accepted now that they emerged from freshwater, multi-celled algae. The embryophytes are informally called land plants because they live primarily in terrestrial habitats (with exceptional members who evolved to live once again in aquatic habitats), while the related green algae are primarily aquatic. Embryophytes are complex multicellular eukaryotes with specialized reproductive organs. The name derives from their innovative characteristic of nurturing the young ...
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Hydrophobe
In chemistry, hydrophobicity is the physical property of a molecule that is seemingly repelled from a mass of water (known as a hydrophobe). In contrast, hydrophiles are attracted to water. Hydrophobic molecules tend to be nonpolar and, thus, prefer other neutral molecules and nonpolar solvents. Because water molecules are polar, hydrophobes do not dissolve well among them. Hydrophobic molecules in water often cluster together, forming micelles. Water on hydrophobic surfaces will exhibit a high contact angle. Examples of hydrophobic molecules include the alkanes, oils, fats, and greasy substances in general. Hydrophobic materials are used for oil removal from water, the management of oil spills, and chemical separation processes to remove non-polar substances from polar compounds. Hydrophobic is often used interchangeably with lipophilic, "fat-loving". However, the two terms are not synonymous. While hydrophobic substances are usually lipophilic, there are exceptions, suc ...
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Casparian Strip
The Casparian strip is a band-like thickening in the center of the root endodermis (radial and tangential walls of endodermal cells) of vascular plants (Pteridophytes and Spermatophytes). The composition of the region is mainly suberin, lignin and some structural proteins, which are capable of reducing the diffusive apoplastic flow of water and solutes into the stele and its width varies between species. The Casparian strip is impervious to water so can control the transportation of water and inorganic salts between the cortex and the vascular bundle, preventing water and inorganic salts from being transported to the stele through the apoplast, so that it must enter the cell membrane and move to the stele through the symplastic pathway, blocking the internal and external objects of the cell. The function and function of mass transportation are similar to that of animal tissues.. The development of the Casparian strip is regulated by transcription factors such as SHORT-ROOT (SHR), ...
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Apoplast
Inside a plant, the apoplast can mean the space outside of cell membranes, where material can diffuse freely; that is, the extracellular spaces. ''Apoplast '' can also refer especially to the continuum of cell walls of adjacent cells; fluid and material flows occurring there or in any extacellular space are called ''apoplastic'' flow or apoplastic transport. The apoplastic route is one way by which water and solutes are transported and distributed to different places through tissues and organs; another way is symplastic flow. To prevent uncontrolled leakage to unwanted places, in certain areas there are barriers to the apoplastic flow: in roots the Casparian strip has this function larification needed On the outside of the skin of certain plant parts is a protective waxy film called plant cuticle to achieve this (protection against e.g. drying out, but also waterproofing against soaking). Air bubbles occupying extracellular spaces can also hinder apoplastic transport. The a ...
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Stele (biology)
In a vascular plant, the stele is the central part of the root or stem containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue (pith) and a pericycle, which, if present, defines the outermost boundary of the stele. Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele. Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs. Protostele The earliest vascular plants had stems with a central core of va ...
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Seedless Plants
A cryptogam (scientific name Cryptogamae) is a plant (in the wide sense of the word) or a plant-like organism that reproduces by spores, without flowers or seeds. The name ''Cryptogamae'' () means "hidden reproduction", referring to the fact that no seed is produced, thus cryptogams represent the non-seed bearing plants. Other names, such as "thallophytes", "lower plants", and "spore plants" are also occasionally used. As a group, Cryptogamae are the opposite of the Phanerogamae () or Spermatophyta (), the seed plants. The best-known groups of cryptogams are algae, lichens, mosses, and ferns, but it also includes non-photosynthetic organisms traditionally classified as plants, such as fungi, slime molds, and bacteria. The classification is now deprecated in Linnaean taxonomy. At one time, the cryptogams were formally recognised as a group within the plant kingdom. In his system for classification of all known plants and animals, Carl Linnaeus (1707–1778) divided the plant k ...
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Suberin
Suberin, cutin and lignins are complex, higher plant epidermis and periderm cell-wall macromolecules, forming a protective barrier. Suberin, a complex polyester biopolymer, is lipophilic, and composed of long chain fatty acids called suberin acids, and glycerol. Suberins and lignins are considered covalently linked to lipids and carbohydrates, respectively, and lignin is covalently linked to suberin, and to a lesser extent, to cutin. Suberin is a major constituent of cork, and is named after the cork oak, ''Quercus suber''. Its main function is as a barrier to movement of water and solutes. Anatomy and physiology Suberin is highly hydrophobic and a somewhat 'rubbery' material. In roots, suberin is deposited in the radial and transverse/tangential cell walls of the endodermal cells. This structure, known as the Casparian strip or Casparian band, functions to prevent water and nutrients taken up by the root from entering the stele through the apoplast. Instead, water must bypass th ...
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Exodermis
The exodermis is a physiological barrier that has a role in root function and protection.  The exodermis is a membrane of variable permeability responsible for the radial flow of water, ions, and nutrients. It is the outer layer of a plant's cortex. The exodermis serves a double function as it can protect the root from invasion by foreign pathogens and ensures that the plant does not lose too much water through diffusion through the root system and can properly replenish its stores at an appropriate rate. Overview and function The exodermis is a specialized type of hypodermis that develops Casparian strips in its cell wall, as well as further wall modifications. The Casparian strip is a band of hydrophobic, corky-like tissue that is found on the outside of the endodermis and the exodermis. Its main function is to prevent solution backflow into the cortex and to maintain root pressure. It is also involved in ensuring that soil is not pulled directly into the root system duri ...
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Epidermis
The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water released from the body into the atmosphere through transepidermal water loss. The epidermis is composed of multiple layers of flattened cells that overlie a base layer (stratum basale) composed of columnar cells arranged perpendicularly. The layers of cells develop from stem cells in the basal layer. The human epidermis is a familiar example of epithelium, particularly a stratified squamous epithelium. The word epidermis is derived through Latin , itself and . Something related to or part of the epidermis is termed epidermal. Structure Cellular components The epidermis primarily consists of keratinocytes ( proliferating basal and differentiated suprabasal), which comprise 90% of its cells, but also contains melanocytes, Langerhans ...
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