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Elmisaurinae
Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that ''Chirostenotes pergracilis'', originally named on the basis of a pair of hands, and ''Citipes elegans'', originally thought to be an ornithomimid, named from a foot, were caenagnathids as well. Anatomy Overall, the anatomy of the caenagnathids is similar to that of the closely related Oviraptoridae, but there are a n ...
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Hagryphus
''Hagryphus'' (meaning " Ha's griffin") is a monospecific genus of caenagnathid dinosaur from southern Utah that lived during the Late Cretaceous (upper Campanian stage, 75.95 Ma) in what is now the Kaiparowits Formation of the Grand Staircase–Escalante National Monument. The type and only species, ''Hagryphus giganteus'', is known only from an incomplete but articulated left manus and the distal portion of the left radius. It was named in 2005 by Lindsay E. Zanno and Scott D. Sampson. ''Hagryphus'' has an estimated length of 2.4–3 metres (8–10 feet) and weight of 50 kilograms (110 lbs). Discovery and naming To date, only a single species of ''Hagryphus'' has been named in 2005 by Lindsay Zanno and Scott Sampson, the type species ''Hagryphus giganteus''. The generic name is derived from Egyptian ''Ha'', the name of the god of the western desert and a Latinised Greek γρύψ (''gryps'') meaning 'griffin' (a mythological bird-like creature). The specific name means "giga ...
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Gigantoraptor
''Gigantoraptor'' () is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005. ''Gigantoraptor'' was the largest oviraptorosaur, reaching in length and in body mass. It had an extensively pneumatized vertebral column and elongated arms and legs. Both femur and tibia measured over in length, an unusual trait among giant theropods. The lower jaws were toothless and ended in a keratinous beak, as seen in other oviraptorosaurs. Though several oviraptorosaur species are known to have developed a full coat of feathers, ''Gigantoraptor'', due to its size, could have lost some of this integument. The genus is classified as an oviraptorosaurian dinosaur, a group of generally small feathered animals. Though it was originally found to represent a basal oviraptorid, subsequent analyses have shown it to be a caenagnathid. It was a giant, ground ...
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Nomingia
''Nomingia'' is a genus of oviraptorid theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia. Discovery and naming The remains, consisting of most of the vertebral column, pelvic girdle and left tibio-tarsus, holotype GIN 100/119, were found in 1994 in layers of the Nemegt Svita, dating to the Maastrichtian. They were named and described as the type species ''Nomingia gobiensis'' by Rinchen Barsbold, Barsbold, Halszka Osmólska, Mahito Watabe, Philip J. Currie, Philip Currie and Khishigjaw Tsogtbaatar in 2000 in paleontology, 2000. The etymology of the Binomial nomenclature, binomial refers to the location where the fossils were found, with the generic name mentioning the Nomingiin Gobi, a nearby part of the Gobi Desert, which is itself mentioned in the specific name (zoology), specific descriptor. A 2021 article by Funston and colleagues suggested ''Nomingia'' is a synonym of ''Elmisaurus''. Description ''Nomingia'' is a medium-sized oviraptoro ...
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Leptorhynchos (dinosaur)
''Leptorhynchos'' (meaning "slender beak") is an extinct genus of caenagnathid dinosaurs known from the Late Cretaceous (Campanian-Maastrichtian aged) Aguja Formation of west Texas United States. It lived about 80.5–72 million years ago. It is distinguished from its relatives ''Chirostenotes'' and ''Anzu'' by its smaller size, and by a more strongly upturned mandible, similar to that of oviraptorids. The specializations of the beak in ''Leptorhynchos'' and other caenagnathids suggest that they were herbivores. The species ''L. elegans'' has since been transferred to the genus ''Citipes'', leaving only the type species ''L. gaddisi'' in the genus. ''Leptorhynchos'' has been placed in the Elmisaurinae. See also * Timeline of oviraptorosaur research This timeline of oviraptorosaur research is a chronological listing of events in the history of paleontology focused on the oviraptorosaurs, a group of beaked, bird-like theropod dinosaurs. The early history of oviraptorosau ...
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Chirostenotes
''Chirostenotes'' ( ; named from Greek 'narrow-handed') is a genus of oviraptorosaurian dinosaur from the late Cretaceous (about 76.5 million years ago) of Alberta, Canada. The type species is ''Chirostenotes pergracilis''. History of discovery ''Chirostenotes'' has a confusing history of discovery and naming. The first fossils of ''Chirostenotes'', a pair of hands, were in 1914 found by George Fryer Sternberg near Little Sandhill Creek in the Campanian Dinosaur Park Formation of Canada, which has yielded the most dinosaurs of any Canadian formation. The specimens were studied by Lawrence Morris Lambe who, however, died before being able to formally name them. In 1924, Charles Whitney Gilmore adopted the name he found in Lambe's notes and described and named the type species ''Chirostenotes pergracilis''. The generic name is derived from Greek ''cheir'', "hand", and ''stenotes'', "narrowness". The specific name means "throughout", ''per~'', "gracile", ''gracilis'', in Latin ...
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Halszka Osmólska
Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Sciences of the University of Poznań before moving to Warsaw and studying at the Warsaw University, which she graduated from in 1955. Since then she worked at the Institute of Paleobiology of the Polish Academy of Sciences (PAN). Between 1983–1988, she served as the institute's director. She was a member of the Polish–Mongolian expeditions to the Gobi Desert (1963–1965 and 1967–1971) and she described many finds from these rocks, often with Teresa Maryańska. Among the dinosaurs she described are: * ''Elmisaurus'' (and Elmisauridae) (1981), ''Hulsanpes'' (1982), '' Borogovia'' (1987), and ''Bagaraatan'' (1996) * with Maryańska, ''Homalocephale'', ''Prenocephale'', and ''Tylocephale'' (and Pachycephalosauria)(1974), ''Bagaceratops'' ...
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Maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. '' Ornitholestes'' and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see '' Eshanosaurus''), and survive today as living birds. Description Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semi-lunate) bone in the wrist (carpus). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characters relating to specific details of the skeleton. Unlike most other saurischian dinosaurs, which h ...
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Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a characteri ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Citipes
''Citipes'' (meaning "fleet-footed") is an extinct genus of caenagnathid dinosaurs known from the Late Cretaceous (Campanian aged) Dinosaur Park of southern Alberta, Canada. It lived about 76.9–75.8 million years ago. The specializations of the beak in ''Citipes'' and other caenagnathids suggest that they were herbivores. The type species, ''C. elegans'' had been previously placed within the genera ''Chirostenotes'', ''Elmisaurus'', '' Leptorhynchos'' and ''Ornithomimus''. History The holotype material, found in 1926 at the Little Sandhill Creek, was originally thought to belong to an ornithomimid; William Arthur Parks William Arthur Parks (11 December 1868 – 3 October 1936) was a Canadian geologist and paleontologist, following in the tradition of Lawrence Lambe. Parks was born in Hamilton, Ontario. After graduating with a Bachelor of Arts in 1892, Parks ... in 1933 assigned it to a new species of ''Ornithomimus'': ''O. elegans''.Parks, W.A., (1933), "New species ...
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Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. Some researchers such as Maryanska ''et al'' (2002) and Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) ...
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