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Eldeceeon
''Eldeceeon'' is an extinct genus of reptiliomorph from the Mississippian (early Carboniferous) of Scotland. It is known from two fossil specimens found within the Viséan-age East Kirkton Quarry in West Lothian. The type and only species, ''E. rolfei'', was named in 1994. ''Eldeceeon'' is thought to be closely related to embolomeres, but it has several distinguishing features including long limbs and a short trunk. Initially known from two crushed partial skeletons, additional specimens have been reported by Ruta & Clack (2006). Description In the East Kirkton Quarry, fossils of ''Eldeceeon'' were found alongside another reptiliomorph called ''Silvanerpeton''. These genera are closely related to each other but represent an unusual group of reptiliomorphs that cannot be placed in any of the major reptiliomorph clades, but may be related to the earliest embolomeres. The shapes of some bones in their skulls Unlike most embolomeres, which could grow over a meter long, ''Eldece ...
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Reptiliomorph
Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians ( lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes ''Homo sapiens'', but not ''Ascaphus truei'' (tailed frog). Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing ''Homo sapiens'', but not ''Pipa pipa'', '' Caecilia tentaculata'', and '' Siren lacertina''. The informal variant of the name, "reptiliomorphs", is also occasionally used to refer to stem-amniotes, i.e. a grade of reptile-like tetrapods that are more closely related to amniotes than they are to lissamphibians, but are not amniotes themselves; the name is used in this meaning e.g. by Ruta, Coates and Quicke (2003). An alternative name, "Anthracosauria", is also commonly used for the group, but is confusing ...
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East Kirkton Quarry
East Kirkton Quarry is a former limestone quarry in West Lothian, Scotland (East Kirkton Limestone), now better known as a fossil site known for terrestrial fossils from the fossil-poor Romer's gap, a 15 million year period at the beginning of the Carboniferous. The rocks and fossils are of Visean age, about 335 million years old. Best known are the labyrinthodont fossils, as the period coincides with the time where the modern lineages of tetrapods are thought to have evolved. Location The quarry is located in the town of Bathgate in West Lothian, Scotland. Geologically, it sits fairly central to the middle of the fossil-rich Scottish Central Lowlands. The site is dominated by volcanic tuff and limestone, and layered silica deposits, indicating the presence of a hot spring associated with volcanism. Benton, M. (2005): Vertebrate Palaeontology 3rd edition. Blackwell Publishing The land next to the quarry itself is developed for housing. Clack, J.A. (2002): Gaining ground: the or ...
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Mississippian (geology)
The Mississippian ( , also known as Lower Carboniferous or Early Carboniferous) is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earlier of two subperiods of the Carboniferous period lasting from roughly 358.9 to 323.2 million years ago. As with most other geochronologic units, the rock beds that define the Mississippian are well identified, but the exact start and end dates are uncertain by a few million years. The Mississippian is so named because rocks with this age are exposed in the Mississippi Valley. The Mississippian was a period of marine transgression in the Northern Hemisphere: the sea level was so high that only the Fennoscandian Shield and the Laurentian Shield were dry land. The cratons were surrounded by extensive delta systems and lagoons, and carbonate sedimentation on the surrounding continental platforms, covered by shallow seas. In North America, where the interval consists primarily of marine limestones, it is treate ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Proterogyrinus
''Proterogyrinus'' is an extinct genus of early tetrapods from the order Embolomeri. Fossil remains of ''Proterogyrinus'' have been found in Scotland, UK, and West Virginia, United States, and date back to the Serpukhovian (mid-Carboniferous period), which is from about 331 to 323 million years ago. The genus was originally named by renowned vertebrate paleontologist Alfred Sherwood Romer in 1970. A comprehensive redescription was later published by Canadian paleontologist Robert Holmes in 1984. The generic name "''Proterogyrinus''" is Greek for "earlier wanderer" or "earlier tadpole". This name was chosen by Romer in keeping with a trend of naming long-bodied early tetrapods (such as ''Eogyrinus'' and ''Crassigyrinus'') with the suffix "-''gyrinus''". Romer hesitated from designating ''Proterogyrinus'' as a true embolomere, because its intercentra (the forward portion of each vertebra) were smaller than its pleurocentra (the rear portion). He used the group Anthracosauria to enc ...
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Pelvic Girdles
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The pelvic region of the trunk includes the bony pelvis, the pelvic cavity (the space enclosed by the bony pelvis), the pelvic floor, below the pelvic cavity, and the perineum, below the pelvic floor. The pelvic skeleton is formed in the area of the back, by the sacrum and the coccyx and anteriorly and to the left and right sides, by a pair of hip bones. The two hip bones connect the spine with the lower limbs. They are attached to the sacrum posteriorly, connected to each other anteriorly, and joined with the two femurs at the hip joints. The gap enclosed by the bony pelvis, called the pelvic cavity, is the section of the body underneath the abdomen and mainly consists of the reproductive organs (sex organs) and the rectum, while the pelvic fl ...
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Pectoral Girdle
The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists of the clavicle, scapula, and coracoid. Some mammalian species (such as the dog and the horse) have only the scapula. The pectoral girdles are to the upper limbs as the pelvic girdle is to the lower limbs; the girdles are the parts of the appendicular skeleton that anchor the appendages to the axial skeleton. In humans, the only true anatomical joints between the shoulder girdle and the axial skeleton are the sternoclavicular joints on each side. No anatomical joint exists between each scapula and the rib cage; instead the muscular connection or physiological joint between the two permits great mobility of the shoulder girdle compared to the compact pelvic girdle; because the upper limb is not usually involved in weight bearing, its stabilit ...
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Eoherpeton
''Eoherpeton'' is the only genus of the family Eoherpetontidae in the extinct suborder Embolomeri. It is known from the Visean and Namurian (now Serpukhovian and lower Bashkirian) stages of the Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ... of Scotland. References External linksThe skull of ''Eoherpeton'', from the original descriptionA 3D reconstruction of the skull, using microCT scans, published by Liz Martin-Silverstone's twitter account
Embolomeres< ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Embolomere
Embolomeri is an Order (biology), order of Tetrapod, tetrapods or Stem-group, stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolution, evolved in the Early Carboniferous (Mississippian age, Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian (geology), Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their Vertebra, vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape. Embolomeres were among the earliest large carnivorous tetrapods, with members such as the crocodilian-like ''Proterogyrinus'' appearing in the Viséan, Visean stage of the Carboniferous. They declined in diversity d ...
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