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Eionaletherium
''Eionaletherium'' is an extinct genus of ground sloth from the Late Miocene coasts of Venezuela containing one species: ''E. tanycnemius''. Taxonomy ''Eionaletherium'' was discovered in the Late Miocene Urumaco Formation in the Falcón State of Venezuela and described in 2015 as a species of mylodontid ground sloth. It is a member of the family Mylodontidae in a clade including ''Bolivartherium'', and ''Glossotherium''. The holotype specimen, IVIC-P-2870, housed at the Venezuelan Institute for Scientific Research, comprises the femurs, a tibia, fibula, some vertebrae, and fragments of the scapulae, ribs, and the ankle bone. The genus name ''Eionaletherium'' comes from the Greek words ''eion'' (shore), ''ale'' (wanderer), and ''therium'' (beast), in reference to the nearshore environment it probably inhabited. The species name ''tanycnemius'' comes from Greek ''tany'' (long) and ''cnemius'' (shin). Description ''Eionaletherium'', based on a femur-to-body-size ratio, is estimat ...
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Thalassocnus
''Thalassocnus'' is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—''T. antiquus'', ''T. natans'', ''T. littoralis'', ''T. carolomartini'', and ''T. yuacensis''—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Horcón formations of Chile. Thalassocninae has been placed in both the families Megatheriidae and Nothrotheriidae. ''Thalassocnus'' evolved several marine adaptations over the course of 4 million years, such as dense and heavy bones to counteract buoyancy, the internal nostrils migrating farther into the head to help with breathing while completely submerged, ...
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Urumaco Formation
The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location The Urumaco formation is located in the Urumaco region in the Caribbean coastal Falcón state. The deposits date from 10 to 5.3 million years ago and the Urumaco formation was deposited in an area with large rivers, swamps, estuaries, lagoons and shallow coastal seas. These conditions in the Late Miocene contrast strongly with the current dry environment in the area today. Fauna Cartilaginous fish There are 21 known species of cartilaginous fishes from the Urumaco Formation, belonging to the orders Lamniformes, Carcharhiniformes, Myliobatiformes and Rajiformes. ''Carcharhinus caquetius'' is an endemic species of predator shark from Urumaco. A large number of well-preserved fossils of the sawfish ''Pristis rostra'' have been found in th ...
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Mylodontidae
Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters and sloths (which includes the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The b ..., living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another family of extinct ground sloths, Scelidotheriidae, as well as to the extant arboreal two-toed sloths, family Choloepus, Choloepodidae; together these make up the superfamily Mylodontoidea. Phylogenetic analyses based on morphology uncovered the relationship between Mylodontidae and Scelidotheriidae; in fact, the latter was for a time considered a subfamily of mylodontids. However, molecular sequence comparisons were needed for the correct placement of Choloepodidae. These studies have been car ...
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Bolivartherium
''Bolivartherium'' is an extinct genus of mylodontine mylodontid sloth that lived during the Late Miocene and Late Pliocene in what is now Venezuela. Fossils have been found in the Codore and Urumaco Formations of Venezuela. Etymology The generic name, ''Bolivartherium'', is named in honour of Libertador Simón Bolívar, a Venezuelan military and political leader. The specific name is derived from the Urumaco Formation in which it was found in. A second species, ''B. codorensis'', was named in 2006 after the Codore Formation in which it was found in. Description ''Bolivartherium'' is a medium-sized mylodontine that was smaller than the quaternary species of ''Lestodon''. It can be distinguished from the latter in having a lower rostrum and the upper caniform which is more curved than in ''Lestodon'', much like ''Lestodon'' sp. from the Monte Hermoso Formation (Montehermosan) of Argentina. The diastema in front of the molariforms is elevated with respect to the occlusal pl ...
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Vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch ...
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Scapula
The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side of the body being roughly a mirror image of the other. The name derives from the Classical Latin word for trowel or small shovel, which it was thought to resemble. In compound terms, the prefix omo- is used for the shoulder blade in medical terminology. This prefix is derived from ὦμος (ōmos), the Ancient Greek word for shoulder, and is cognate with the Latin , which in Latin signifies either the shoulder or the upper arm bone. The scapula forms the back of the shoulder girdle. In humans, it is a flat bone, roughly triangular in shape, placed on a posterolateral aspect of the thoracic cage. Structure The scapula is a thick, flat bone lying on the thoracic wall that provides an attachment for three groups of muscles: intrin ...
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Ankle Bone
The talus (; Latin for ankle or ankle bone), talus bone, astragalus (), or ankle bone is one of the group of Foot#Structure, foot bones known as the tarsus (skeleton), tarsus. The tarsus forms the lower part of the ankle joint. It transmits the entire weight of the body from the lower legs to the foot.Platzer (2004), p 216 The talus has joints with the two bones of the lower leg, the tibia and thinner fibula. These leg bones have two prominences (the Lateral malleolus, lateral and Medial malleolus, medial malleoli) that articulation (anatomy), articulate with the talus. At the foot end, within the tarsus, the talus articulates with the calcaneus (heel bone) below, and with the curved navicular bone in front; together, these foot articulations form the Ball-and-socket joint, ball-and-socket-shaped talocalcaneonavicular joint. The talus is the second largest of the Tarsus (skeleton), tarsal bones; it is also one of the bones in the human body with the highest percentage of its surfa ...
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Ancient
Ancient history is a time period from the History of writing, beginning of writing and recorded human history to as far as late antiquity. The span of recorded history is roughly 5,000 years, beginning with the Sumerian language, Sumerian cuneiform script. Ancient history covers all continents inhabited by humans in the period 3000 BCAD 500. The three-age system periodizes ancient history into the Stone Age, the Bronze Age, and the Iron Age, with recorded history generally considered to begin with the Bronze Age. The start and end of the three ages varies between world regions. In many regions the Bronze Age is generally considered to begin a few centuries prior to 3000 BC, while the end of the Iron Age varies from the early first millennium BC in some regions to the late first millennium AD in others. During the time period of ancient history, the world population was already Exponential growth, exponentially increasing due to the Neolithic Revolution, which was in full pro ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Littoral Zone
The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas that are permanently submerged — known as the ''foreshore'' — and the terms are often used interchangeably. However, the geographical meaning of ''littoral zone'' extends well beyond the intertidal zone to include all neritic waters within the bounds of continental shelves. Etymology The word ''littoral'' may be used both as a noun and as an adjective. It derives from the Latin noun ''litus, litoris'', meaning "shore". (The doubled ''t'' is a late-medieval innovation, and the word is sometimes seen in the more classical-looking spelling ''litoral''.) Description The term has no single definition. What is regarded as the full extent of the littoral zone, and the way the littoral zone is divided into subregions, varies in different c ...
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Tibia
The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects the knee with the ankle. The tibia is found on the medial side of the leg next to the fibula and closer to the median plane. The tibia is connected to the fibula by the interosseous membrane of leg, forming a type of fibrous joint called a syndesmosis with very little movement. The tibia is named for the flute ''tibia''. It is the second largest bone in the human body, after the femur. The leg bones are the strongest long bones as they support the rest of the body. Structure In human anatomy, the tibia is the second largest bone next to the femur. As in other vertebrates the tibia is one of two bones in the lower leg, the other being the fibula, and is a component of the knee and ankle joints. The ossification or formation of the bone ...
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Pachyosteosclerosis
Pachyosteosclerosis is a combination of thickening (pachyostosis) and densification (osteosclerosis) of bones. It makes bones more heavy, but also more fragile. The condition often occurs in aquatic vertebrates, especially those living in shallow waters,Houssaye, A. (2009). "Pachyostosis" in aquatic amniotes: a review. Integrative Zoology 4(4): 325-340. creating ballast as an adaptation for maintaining neutral buoyancy and horizontal trim. It is in no way pathological. To resist bend, it frequently is found especially in ventral bones, whereas concentration near the lungs helps in maintaining trim. Examples of animals showing pachyosteosclerosis are seacows (dugongs and manatees), the extinct Plesiosauria and Mesosauria and extinct aquatic sloths.Amson, E., C. de Muizon, M. Laurin, C. Argot and V. de Buffrenil (2014). Gradual adaptation of bone structure to aquatic lifestyle in extinct sloths from Peru. Proceedings of the Royal Society B-Biological Sciences 281. See also * Pachyo ...
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