Danziella
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Danziella
''Danziella'' is a genus of extinct vascular plants of the Early Devonian (around ). Fossils found in the Artois region of northern France were first described as ''Zosterophyllum artesianum'', but a later review by Edwards showed that they did not fit the circumscription of that genus. Description Compressed fossils were found in the Artois region of northern France, in rocks which were originally thought to be of Pragian age (around ) but more recently have been considered to be probably of Emsian origin (around ). Plants consisted of smooth leafless stems (axes) up to 1.5 mm wide and were at least 85 mm high. They branched at right angles. Spore-forming organs or sporangia were borne on all sides, spaced irregularly on stalks up to 3 mm long which held them horizontally, and not forming a distinct 'spike'. Individual sporangia were elliptical in shape, around 3 mm wide by 2 mm high, splitting into two 'valves' along a line opposite to the stalks in or ...
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Zosterophylls
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Zosterophyllum
''Zosterophyllum'' was a genus of Silurian-Devonian vascular land plants with naked branching axes on which usually kidney-shaped sporangia were arranged in lateral positions. It is the type genus for the group known as zosterophylls, thought to be part of the lineage from which modern lycophytes evolved. More than 20 species have been described. Description The diagnostic features of the genus have changed since its first description in 1892, as the original species (''Zosterophyllum myretonianum'') has become better known, and as other species have been discovered. ''Zosterophyllum'' is a vascular plant. The axes (stems) are naked, lacking leaves or outgrowths ("enations"). When branching occurs, the branches are either isotomous (equally sized) or pseudomonopodial (one branch is larger than the other but still clearly involves division of the original axis rather a distinct side growth). The sporangia are upright on short stalks. In face view, they are flattened, usually kid ...
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Zosterophyll
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Artois
Artois ( ; ; nl, Artesië; English adjective: ''Artesian'') is a region of northern France. Its territory covers an area of about 4,000 km2 and it has a population of about one million. Its principal cities are Arras (Dutch: ''Atrecht''), Saint-Omer, Lens, and Béthune. It is the eponym for the term '' artesian''. Location Artois occupies the interior of the Pas-de-Calais ''département'',"Artois" in ''The New Encyclopædia Britannica''. Chicago: Encyclopædia Britannica Inc., 15th ed., 1992, Vol. 1, p. 607. the western part of which constitutes the former Boulonnais. Artois roughly corresponds to the arrondissements of Arras, Béthune, Saint Omer, and Lens, and the eastern part of the arrondissement of Montreuil. It occupies the western end of the coalfield which stretches eastward through the neighbouring Nord ''département'' and across central Belgium. History Originally a feudal county itself, Artois was annexed by the county of Flanders. It came to France in ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

{{Geological history Early Devonian, Geological epochs Devonian geochronology, *01 ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics ...
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Pragian
The Pragian is one of three faunal stages in the Early Devonian Epoch. It lasted from 410.8 ± 2.8 million years ago to 407.6 ± 2.8 million years ago. It was preceded by the Lochkovian Stage and followed by the Emsian Stage. The most important Lagerstätte of the Pragian is Rhynie chert in Scotland. It is named after the city of Prague. The GSSP is located within the Prague Formation at Velká Chuchle, Prague. In North America the Pragian Stage is represented by Siegenian or Deerparkian time. Pragian life The first ammonoids (order Agoniatitida) appeared in later parts of this stage (at the lower boundary of the Zlichovian stage as it was known in Siberian representations). They were descended from bactritoid nautiloid Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species ... ances ...
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Emsian
The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the Ems river in Germany. The GSSP is located in the Zinzil'ban Gorge in the Kitab State Geological Reserve of Uzbekistan, above the contact with the Madmon Formation. In North America the Emsian Stage is represented by Sawkill or Sawkillian time. Biological events During this period, earliest known agoniatitid ammonoid fossils began appearing within this stage after first appearing in previous stage and began to evolutionarily radiate within this stage, in which a new ammonoid order Goniatitida rises in the end of Zlichovian stage (Siberian representation; corresponds to early Eifelian and after the end of Early Devonian, before 391.9 mya). Later agoniatitid ammonoids would die out in the Taghanic event in the upper middle Givetian ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Vascular Plant
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the term ...
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Cooksonia (plant)
''Cooksonia'' is an extinct group of primitive land plants, treated as a genus, although probably not monophyletic. The earliest ''Cooksonia'' date from the middle of the Silurian (the Wenlock epoch); the group continued to be an important component of the flora until the end of the Early Devonian, a total time span of . While ''Cooksonia'' fossils are distributed globally, most type specimens come from Britain, where they were first discovered in 1937. ''Cooksonia'' includes the oldest known plant to have a stem with vascular tissue and is thus a transitional form between the primitive non-vascular bryophytes and the vascular plants. Description Only the sporophyte phase of ''Cooksonia'' is currently known (i.e. the phase which produces spores rather than gametes). Individuals were small, a few centimetres tall, and had a simple structure. They lacked leaves, flowers and roots—although it has been speculated that they grew from a rhizome that has not been preserved. They had ...
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Renalia
''Renalia'' is a genus of extinct vascular plants from the Early Devonian (around ). It was first described in 1976 from compressed fossils in the Battery Point Formation (Gaspé, Québec, Canada). It is difficult to reconstruct the original form of the complete plant, but it appears to have consisted of leafless branching stems whose side branches had sporangia (spore-forming organs) at their tips. It is regarded as an early relative of the lycophytes (clubmosses and relatives). Description The species ''R. hueberi'' was described by Patricia G. Gensel in 1976 from compressed fossils. As with all compressions, it is difficult to reconstruct the entire plant, but the sporophyte is thought to have been about 30 cm tall, with bare stems (axes) up to 1.5 mm wide. The main stem had lateral branches which divided into two equal branches (dichotomized) several times. Tracheids (specialized cells serving the purpose of water transport) were present in the stems. The rounde ...
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