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Dolichophonus
''Dolichophonus'' is an extinct genus of scorpion known from the Silurian aged Gutterford Burn Eurypterid bed, in the Pentland Hills in Scotland. It is currently considered the world's oldest known scorpion, as the older '' Parioscorpio'' named in 2020 has been found to be an unrelated arthropod. lt is sometimes referred to '' Palaeophonus''. Discovery The specimen of ''Dolichophonus'' was collected by Mr Hardie of Bavelaw Castle. When he died his collection was given to the Edinburgh Museum of Science and Art (now National Museum of Scotland), where it was then studied by Malcolm Laurie. He named seven new species of Eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is l ...s from the collection, along with this scorpion, which he named. References Prehistoric scor ...
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Scorpion
Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always ending with a stinger. The evolutionary history of scorpions goes back 435 million years. They mainly live in deserts but have adapted to a wide range of environmental conditions, and can be found on all continents except Antarctica. There are over 2,500 described species, with 22 extant (living) families recognized to date. Their taxonomy is being revised to account for 21st-century genomic studies. Scorpions primarily prey on insects and other invertebrates, but some species hunt vertebrates. They use their pincers to restrain and kill prey, or to prevent their own predation. The venomous sting is used for offense and defense. During courtship, the male and female grasp each other's pincers and dance while he tries to move her onto ...
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Chelicerata
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all cover ...
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Parioscorpio
''Parioscorpio'' is an extinct genus of arthropod containing the species ''P. venator'' known from the Silurian-aged Waukesha Biota of the Brandon Bridge Formation near Waukesha, Wisconsin. Taxonomy The fossils were originally discovered in 1985, tentatively identified as a branchiopod or remipede crustacean but were neglected for decades. In 2016, some of the fossils now assigned to ''Parioscorpio'' were given the name ''Latromirus'' and were assigned to an extinct group of early Paleozoic arthropods known as cheloniellids in a Ph.D dissertation, but the name was never published in a peer-reviewed journal and is therefore not valid in accordance with the International Code of Zoological Nomenclature. Upon initial publication in 2020, ''Parioscorpio'' was considered the world's oldest and most primitive known scorpion, older than ''Dolichophonus'' from Scotland by several million years. In 2021, the fossils were reanalysed, and ''Parioscorpio'' was found not to be a scorpion, b ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone ...
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Telychian
In the geologic timescale, the Telychian is the age of the Llandovery Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon. The Telychian Age was between 438.5 ± 1.2 million years ago (Ma) and 433.4 ± 0.8 Ma. The Telychian Age succeeds the Aeronian Age and precedes the Sheinwoodian Age. The name of the interval is derived from the Pen-lan-Telych Farm near Llandovery, Powys, Wales. The GSSP is located within the Wormwood Formation. It ended with the Ireviken event. Ireviken event The Ireviken event was the first of three relatively minor extinction events (the Ireviken, Mulde, and Lau events) during the Silurian Period. It occurred at the Llandovery/ Wenlock boundary (mid Silurian, ). The event is best recorded at Ireviken, Gotland, where over 50% of trilobite species became extinct; 80% of the global conodont species also become extinct in this interval. Anatomy of the event The event lasted around 200,000 years, spanning the base of the Wenlo ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is ...
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Arachnid
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Almost all adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 100,000 named species, of which 47,000 are species of spiders. Morphology Almost all adult arachnids have eight legs, unlike adult i ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus '' Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event (or some ...
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Pentland Hills
The Pentland Hills are a range of hills southwest of Edinburgh, Scotland. The range is around in length, and runs southwest from Edinburgh towards Biggar and the upper Clydesdale. Etymology The name is first recorded for the farm of Pentland ( 1050, 1200) and probably derives from Brythonic ''pen llan'', head or top end of the church or enclosure. In the late 15th to mid-16th centuries, land transfers refer to Pentlandmure and Pentland – documents that also list adjacent parcels of land with such still-recognisable names as Loganehous, Hilend, Boghall and Mortounhall. 'Muir', in Pentlandmure, describes common grazings where the farm's livestock would be pastured in summer; and gradually the name was linked more specifically with the slopes of the nearby hills (perhaps Allermuir, Woodhouselee or Castlelaw). The name is completely unrelated to the name of the Pentland Firth in the north of Scotland. Timothy Pont mapped the area in the 1590s, and his work appeared in the ma ...
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