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Divergent Evolution
Divergent evolution or divergent selection is the accumulation of differences between closely related populations within a species, leading to speciation. Divergent evolution is typically exhibited when two populations become separated by a geographic barrier (such as in allopatric or peripatric speciation) and experience different selective pressures that drive adaptations to their new environment. After many generations and continual evolution, the populations become less able to interbreed with one another. The American naturalist J. T. Gulick (1832–1923) was the first to use the term "divergent evolution", with its use becoming widespread in modern evolutionary literature. Classic examples of divergence in nature are the adaptive radiation of the finches of the Galapagos or the coloration differences in populations of a species that live in different habitats such as with pocket mice and fence lizards. The term can also be applied in molecular evolution, such as to pr ...
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Darwin's Finches
Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or tribe Geospizini. They belong to the tanager family and are not closely related to the true finches. The closest known relative of the Galápagos finches is the South American '' Tiaris obscurus''. They were first collected when the second voyage of the ''Beagle'' visited the Galápagos Islands, with Charles Darwin on board as a gentleman naturalist. Apart from the Cocos finch, which is from Cocos Island, the others are found only on the Galápagos Islands. The term "Darwin's finches" was first applied by Percy Lowe in 1936, and popularised in 1947 by David Lack in his book ''Darwin's Finches''. Lack based his analysis on the large collection of museum specimens collected by the 1905–06 Galápagos expedition of the California Academy ...
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Analogous Structures
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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Chronospecies
A chronospecies is a species derived from a anagenesis, sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is physically, Comparative anatomy, morphologically, and/or Genetics, genetically distinct from the original ancestors. Throughout the change, there is only one species in the lineage at any point in time, as opposed to cases where divergent evolution produces contemporary species with a common ancestor. The related term paleospecies (or palaeospecies) indicates an extinct species only identified with fossil material. That identification relies on distinct similarities between the earlier fossil specimens and some proposed descendant although the exact relationship to the later species is not always defined. In particular, the range of variation within all the early fossil specimens does not exceed the observed range that exists in t ...
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Devolution (biology)
Devolution, de-evolution, or backward evolution (not to be confused with dysgenics) is the notion that species can revert to supposedly more primitive forms over time. The concept relates to the idea that evolution has a purpose (teleology) and is progressive (orthogenesis), for example that feet might be better than hooves or lungs than gills. However, evolutionary biology makes no such assumptions, and natural selection shapes adaptations with no foreknowledge of any kind. It is possible for small changes (such as in the frequency of a single gene) to be reversed by chance or selection, but this is no different from the normal course of evolution and as such de-evolution is not compatible with a proper understanding of evolution due to natural selection. In the 19th century, when belief in orthogenesis was widespread, zoologists (such as Ray Lankester and Anton Dohrn) and the palaeontologists Alpheus Hyatt and Carl H. Eigenmann advocated the idea of devolution. The concept app ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Husky
Husky is a general term for a dog used in the polar regions, primarily and specifically for work as sled dogs. It refers to a traditional northern type, notable for its cold-weather tolerance and overall hardiness. Modern racing huskies that maintain arctic breed traits (also known as Alaskan huskies) represent an ever-changing crossbreed of the fastest dogs. Huskies have continued to be used in sled-dog racing, as well as expedition and trek style tour businesses, and as a means of essential transportation in rural communities. Huskies are also kept as pets, and groups work to find new pet homes for retired racing and adventure-trekking dogs. History Nearly all dogs' genetic closeness to the gray wolf is due to admixture. However, several Arctic breeds also show a genetic closeness with the now-extinct Taimyr wolf of North Asia due to admixture: the Siberian Husky and Greenland Dog (which are also historically associated with Arctic human populations) and to a lesser ex ...
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Malamute
The Alaskan Malamute () is a large breed of dog that was originally bred for its strength and endurance to haul heavy freight as a sled dog and hound. It is similar to other arctic breeds such as the husky, the spitz, the Greenland Dog, Canadian Eskimo Dog, the Siberian Husky, and the Samoyed. Lineage Although it is believed that the first dogs arrived in the Americas 12,000 years ago, people and their dogs did not settle in the Arctic until the Paleo-Eskimo people 4,500 years ago, followed by the Thule people 1,000 years ago, with both originating from Siberia. Malamutes were thought to be bred by the Malimiut Inupiaq people of Alaska's Norton Sound region. The Malamute has been identified as a basal breed that predates the emergence of the modern breeds in the 19th century. A study in 2013 showed that the Alaskan Malamute has a similar east Asian origin to, but is not clearly related to, the Greenland Dog and the Canadian Eskimo Dog, but contains a possible admixture ...
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Mitochondrial DNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA is only a small portion of the DNA in a eukaryotic cell; most of the DNA can be found in the cell nucleus and, in plants and algae, also in plastids such as chloroplasts. Human mitochondrial DNA was the first significant part of the human genome to be sequenced. This sequencing revealed that the human mtDNA includes 16,569 base pairs and encodes 13 proteins. Since animal mtDNA evolves faster than nuclear genetic markers, it represents a mainstay of phylogenetics and evolutionary biology. It also permits an examination of the relatedness of populations, and so has become important in anthropology and biogeography. Origin Nuclear and mitochondrial DNA are thought to be of separate evolutionary origin, with the mtDNA being derived ...
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Wolf
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly understood, comprise wild subspecies. The wolf is the largest extant member of the family Canidae. It is also distinguished from other ''Canis'' species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller ''Canis'' species, such as the coyote and the golden jackal, to produce fertile hybrids with them. The banded fur of a wolf is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white. Of all members of the genus ''Canis'', the wolf is most specialized for cooperative game hunting as demonstrated by its physical adaptations to tackling large prey, its more social nature, and its highly advanc ...
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Origin Of The Domestic Dog
The domestication of the dog was the process which created the domestic dog. This included the dog's genetic divergence from the wolf, its domestication, and the emergence of the first dogs. Genetic studies show that all ancient and modern dogs share a common ancestry and descended from an ancient, now-extinct wolf population – or closely related wolf populations – which was distinct from the Wolf, modern wolf lineage. The dog's similarity to the grey wolf is the result of substantial dog-into-wolf gene flow, with the modern grey wolf being the dog's nearest living relative. An extinct Pleistocene wolf, Late Pleistocene wolf may have been the ancestor of the dog. The dog is a member of the Evolution of the wolf#Wolf-like canids, wolf-like canids. The genetic divergence between the dog's ancestor and modern wolves occurred between 40,000 and 30,000 years ago, just before or during the Last Glacial Maximum (20,000–27,000 years ago). This timespan represents the upper time-limi ...
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Darwin's Finches
Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or tribe Geospizini. They belong to the tanager family and are not closely related to the true finches. The closest known relative of the Galápagos finches is the South American '' Tiaris obscurus''. They were first collected when the second voyage of the ''Beagle'' visited the Galápagos Islands, with Charles Darwin on board as a gentleman naturalist. Apart from the Cocos finch, which is from Cocos Island, the others are found only on the Galápagos Islands. The term "Darwin's finches" was first applied by Percy Lowe in 1936, and popularised in 1947 by David Lack in his book ''Darwin's Finches''. Lack based his analysis on the large collection of museum specimens collected by the 1905–06 Galápagos expedition of the California Academy ...
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Parallel Evolution
Parallel evolution is the similar development of a trait in distinct species that are not closely related, but share a similar original trait in response to similar evolutionary pressure.Zhang, J. and Kumar, S. 1997Detection of convergent and parallel evolution at the amino acid sequence level. ''Mol. Biol. Evol.'' 14, 527-36. Parallel vs. convergent evolution Given a particular trait that occurs in each of two lineages descended from a specified ancestor, it is possible in theory to define parallel and convergent evolutionary trends strictly, and distinguish them clearly from one another. However the criteria for defining convergent as opposed to parallel evolution often are unclear in practice, so that arbitrary diagnosis is common in some cases. When two species are similar in a particular character, evolution is defined as parallel if the ancestors shared that similarity; if they did not, the evolution of that character in those species is defined as convergent. However, thi ...
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