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Dissorophid
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be d ...
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Temnospondyli
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bon ...
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Dissorophus
''Dissorophus'' (DI-soh-ROH-fus) (meaning "double roof" for two layers of armor) is an extinct genus of temnospondyl amphibian that lived during the Early Permian Period about 273 million years ago. Its fossils have been found in Texas and in Oklahoma in North America. Its heavy armor and robust build indicate ''Dissorophus'' was active on land, similar to other members of the clade Dissorophidae that are known from the Late Carboniferous to the Early Permian periods. ''Dissorphus'' is distinguished by its small body size, disproportionately large head and short trunk. The American paleontologist Edward Drinker Cope first briefly described ''Dissorophus'' in 1895,Cope. E.D. 1895. A batrachian armadillo. ''American Naturalist'' 29:99/ref> likely deriving the genus name from Ancient Greek δισσός/dissos "double" and ὀροφή/orophe "roof" to refer to the double layer of armor formed by horizontal "spinous branches" at the top of the neural spines of the vertebrae that "tou ...
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Trematopidae
Trematopidae is a family of dissorophoid temnospondyl spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany. History of study The clade Trematopidae was first proposed by American paleontologist S.W. Williston in 1910, although it was named as "Trematopsidae" following the historical (but inaccurate) derivation from the genus "''Trematops''" (now synonymized with ''Acheloma''). British paleontologist D.M.S. Watson proposed a related clade in 1919, Achelomidae, for ''Acheloma'', based on perceived differences separating the taxa; this is now considered a junior synonym of Trematopidae following guidelines of historical precedent. 19th century history In 1882, American paleontologist Edward Drinker Cope na ...
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Aspidosaurus
''Aspidosaurus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. Description Like other dissorophids, ''Aspidosaurus'' species had a single row of plates formed by expansions of the neural spines.A Description of ''Aspidosaurus novomexicanus'' Williston
Permo-Carboniferous Vertebrates from New Mexico, p7-11. Retrieved 2011-09-11.


Taxonomy

In 1911, Paul Miller discovered the remains of various dissorophid bones in that were attributed to a new species, ''Aspidosaurus novomexicanus''. The skull closely resembled that of a specimen of ''

Broiliellus
''Broiliellus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. ''Broiliellus'' is most closely related to the genus ''Dissorophus'', and both have been placed in the subfamily Dissorophinae. ''Broiliellus'' is known from five species from the Early Permian: the type species is ''Broiliellus texensis,'' and the other species are ''Broiliellus brevis,'' ''Broiliellus olsoni, Broiliellus arroyoensis,'' and ''Broiliellus reiszi''. An additional species, ''Broiliellus novomexicanus'', which was originally named ''Aspidosaurus novomexicanus'', is now thought to fall outside the genus as a member of the subfamily Eucacopinae. History of study ''Broiliellus'' was first named by American paleontologist S.W. Williston in 1914 based on two nearly complete skulls in articulation with postcranial material from the early Permian of Texas; this species was given the name ''Broiliellus texensis'', the genus name being for the German paleontologist Ferdinand B ...
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Kamacops
''Kamacops'' is a genus of dissorophid temnospondyls known from the Middle to Late Permian of Russia that was described by Yuri Gubin in 1980. It is known from a single species, ''Kamacops acervalis,'' material of which is currently reposited in the Paleontological Institute of the Russian Academy of Sciences. ''Kamacops'' is one of the youngest dissorophids, along with '' Iratusaurus'' and '' Zygosaurus'' from Russia and ''Anakamacops'' from China and was one of the largest known dissorophids, with an estimated skull length of 24–30 cm. It is typically recovered as being most closely related to ''Zygosaurus'' and to the North American ''Cacops ''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids) ...''''.'' A detailed study of the braincase region was performed by Schoch (1999), ...
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Parioxys
''Parioxys'' is an extinct genus of temnospondyl amphibian from the Early Permian of Texas. History of study The type species, ''Parioxys ferricolus'', was named in 1878 by American paleontologist Edward Drinker Cope based on two badly preserved skulls that were collected from the early Permian Texas red-beds. Egyptian paleontologist Youssef S. Moustafa, described new material of ''P. ferricolus'' from other localities in Texas. This taxon was recently redescribed by Schoch & Sues (2022). Moustafa also described another species, ''P. romeri'', on the basis of an isolated humerus, but this was regarded as being indeterminate. A second definitive species, ''P. bolli'', was described by Canadian paleontologist Robert Carroll in 1964. This taxon is only known from postcranial material. Relationships ''Parioxys'' was historically considered to be closely related to eryopoids, more specifically the well-known '' Eryops megalocephalus'', which was collected from the same locality ...
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Brevidorsum
''Brevidorsum'' is an extinct genus of dissorophoidean euskelian temnospondyl within the family Dissorophidae. See also * Prehistoric amphibian * List of prehistoric amphibians This list of prehistoric amphibians is an attempt to create a comprehensive listing of all Genus, genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accep ... References Dissorophids Cisuralian temnospondyls of North America Prehistoric amphibian genera {{temnospondyli-stub ...
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Cacops Aspidephorus
''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids) whose ontogeny is known. ''Cacops'' fossils were almost exclusively known from the Cacops Bone Bed of the Lower Permian Arroyo Formation of Texas for much of the 20th century. New material collected from the Dolese Brothers Quarry, near Richards Spur, Oklahoma in the past few decades has been recovered, painting a clearer picture of what the animal looked and acted like. History of discovery ''Cacops aspidephorus'' is the most famous dissorophid, in part due to a majority of its skeleton having been known for over a century. Over 50 specimens have been found in the ''Cacops'' Bone Bed in Baylor County, Texas, which is now flooded by the dammed Lake Kemp. However, many of the specimens are covered in calcite, which penetrates the bone tis ...
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Conjunctio Multidens
''Conjunctio'' is an extinct genus of dissorophid temnospondyl amphibian from the early Permian of New Mexico. The type species, ''Conjunctio multidens'', was named by paleontologist Robert L. Carroll in 1964. History of study The holotype specimen was found in 1911 in the Lower Permian Abo Formation in New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker .... It consists of the skull and postcranial material including femora, humeri, scapulae, pelvis, a section of the vertebral column, and osteoderms. It was originally described by Case et al. (1913) as a referred specimen of ''Aspidosaurus'' (sometimes "''Broiliellus''") ''novomexicanus'' but was subsequently determined to be a distinct species by Carroll (1964). Carroll also identified a third specimen of ''Conjunctio'', ...
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Astreptorhachis
''Astreptorhachis'' is an extinct genus of Late Carboniferous dissorophoid temnospondyl within the family Dissorophidae. It is known only from one species, ''Astreptorhachis ohioensis'', that was collected from Jefferson County, Ohio by the Ohio Geological Survey in 1953 and described by Peter Vaughn in 1971. The holotype and only known specimen consists of a few neural spines and is currently reposited in the Smithsonian Museum of Natural History. The genus name derives from the Greek words ''astreptos'' ("inflexible") and ''rhachis'' ("backbone"). The specimen was recognized as being similar to the dissorophid '' Platyhystrix rugosus'' from the southwestern United States in have greatly elongate neural spines. ''Astreptorhachis'' is differentiated from ''Platyhystrix'' by the fusion of successive neural spines and the extensively developed tubercles that cover the external surfaces. It is speculated that the elongation of the spines served to stiffen the backbone, being advantage ...
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Iratusaurus
''Iratusaurus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. It was described by Gubin (1980) on the basis of a fragmentary posterior skull. It is estimated to have been comparably large to ''Kamacops'', another Russian dissorophid, but little more can be said about it, and it is rarely mentioned in comparative descriptions and has never been tested in a phylogenetic analysis. Distinguishing features include a large, triangular otic notch and a median crest on the postparietals. See also * List of prehistoric amphibians This list of prehistoric amphibians is an attempt to create a comprehensive listing of all Genus, genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accep ... References Dissorophids Permian temnospondyls Fossils of Russia Prehistoric amphibian genera {{temnospondyli-stub ...
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