Diplocraterion
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Diplocraterion
''Diplocraterion'' is an ichnogenus describing vertical U-shaped burrows having a spreite (weblike construction) between the two limbs of the U. The spreite of an individual ''Diplocraterion'' trace can be either protrusive (between the paired tubes) or retrusive (below the paired tubes). Some ichnospecies have both types (e.g., ''Diplocraterion yoyo''). The presence/absence of funnel-shaped openings should not be used as an ichnotaxobase due to the high probability that the upper portions of the trace may have been eroded away. Observation of the orientation of ''Diplocraterion'' in the field is frequently used to determine the way up of rock strata at outcrop. There are several ichnospecies of ''Diplocraterion''. Ethology The various ichnospecies of ''Diplocraterion'' provide a good example of how ethology (animal-substrate interactions and behavior) can be interpreted from trace fossils. ''Diplocraterion'' is a classic example of equilibrichnia (equilibrium traces). Thes ...
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Diplocraterion Parallelum - Richter1926
''Diplocraterion'' is an ichnogenus describing vertical U-shaped burrows having a spreite (weblike construction) between the two limbs of the U. The spreite of an individual ''Diplocraterion'' trace can be either protrusive (between the paired tubes) or retrusive (below the paired tubes). Some ichnospecies have both types (e.g., ''Diplocraterion yoyo''). The presence/absence of funnel-shaped openings should not be used as an ichnotaxobase due to the high probability that the upper portions of the trace may have been eroded away. Observation of the orientation of ''Diplocraterion'' in the field is frequently used to determine the way up of rock strata at outcrop. There are several ichnospecies of ''Diplocraterion''. Ethology The various ichnospecies of ''Diplocraterion'' provide a good example of how ethology (animal-substrate interactions and behavior) can be interpreted from trace fossils. ''Diplocraterion'' is a classic example of equilibrichnia (equilibrium traces). Thes ...
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Spreite
Spreite, meaning Lamina (leaf), leaf-blade in German (or spreiten, the plural form in German language, German) is a stacked, curved, layered structure that is characteristic of certain trace fossils. They are formed by invertebrate organisms tunneling back and forth through sediment in search of food. The organism moves perpendicularly just enough at the start of each back-and-forth pass so that it avoids reworking a previously tunneled area, thereby ensuring that it only makes feeding passes through fresh, unworked sediment. Two types of spreiten are generally recognized. Protrusive spreiten result from movement of the organism away from its burrow entrance (i.e., a downward movement in vertical burrows), whereas retrusive spreiten result from movement towards the burrow entrance (i.e., an upward movement in vertical burrows). Vertical burrows with retrusive spreiten are also referred to as "escape burrows", as they represent attempts by the organism during periods of high sedime ...
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Tuscarora Formation
The Silurian Tuscarora Formation — also known as Tuscarora Sandstone or Tuscarora Quartzite — is a mapped bedrock unit in Pennsylvania, Maryland, West Virginia, and Virginia, USA. Description The Tuscarora is a thin- to thick-bedded fine-grained to coarse-grained orthoquartzite. It is a white to medium-gray or gray-green subgraywacke, sandstone, siltstone and shale, cross-stratified and conglomeratic conglomerate in parts, containing a few shale interbeds.Berg, T.M., Edmunds, W.E., Geyer, A.R. and others, compilers, (1980). Geologic Map of Pennsylvania: Pennsylvania Geologic Survey, Map 1, scale 1:250,000. Details of the type locality and of stratigraphic nomenclature for this unit as used by the U.S. Geological Survey are available on-line at the National Geologic Map Database. The Tuscarora and its lateral equivalents are the primary ridge-formers of the Ridge-and-Valley Appalachians in the eastern United States It is typically 935 feet thick in Pennsylvania, and in Mar ...
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Burrow Fossils
Burrow fossils are the remains of burrows - holes or tunnels excavated into the ground or seafloor - by animals to create a space suitable for habitation, temporary refuge, or as a byproduct of locomotion preserved in the rock record. Because burrow fossils represent the preserved byproducts of behavior rather than physical remains, they are considered a kind of trace fossil. One common kind of burrow fossil is known as Skolithos, and the similar Trypanites, Ophiomorpha and Diplocraterion. Vertebrate burrows Fish burrows Fossil Lungfish burrows are preserved in the Rocky Point Member of the Chinle Formation in Canyonlands National Park. Invertebrate burrows Examples are Treptichnus pedum ''Treptichnus'' (formerly named ''Phycodes'', ''Manykodes'' by J. Dzik, and also known as ''Trichophycus''See e.gfossiilid.info: paleodiversity in Baltoscandia: Trichophycus pedum/ref>) is the preserved burrow of an animal. As such, it is regar ... and Arenicolites franconicus. Footnotes ...
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Way Up Structure
A way up structure'', ''way up criterion, or geopetal indicator is a characteristic relationship observed in a sedimentary or volcanic rock, or sequence of rocks, that makes it possible to determine whether they are the right way up (i.e. in the attitude in which they were originally deposited, also known as "stratigraphic up" or "younging upwards") or have been overturned by subsequent deformation. This technique is particularly important in areas affected by thrusting and where there is a lack of other indications of the relative ages of beds within the sequence, such as in the Precambrian where fossils are rare. The original definition comes from Bruno Sander in 1936, translated from German to English in 1951, which states: ''Geopetal Fabrics'' - All the widely distributed spatial characters of a fabric that enable us to determine what was the relation of "top" to "bottom" at the time when the rock was formed are termed ''geopetal'' fabrics. Such fabrics are mechanical and che ...
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Trace Fossil
A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization. The study of such trace fossils is ichnology and is the work of ichnologists. Trace fossils may consist of impressions made on or in the substrate by an organism. For example, burrows, borings (bioerosion), urolites (erosion caused by evacuation of liquid wastes), footprints and feeding marks and root cavities may all be trace fossils. The term in its broadest sense also includes the remains of other organic material produced by an organism; for example coprolites (fossilized droppings) or chemical markers (sedimentological structures produced by biological means; for example, the formation of stromatolites). H ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during ...
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Rhizocorallium
''Rhizocorallium'' is an ichnogenus of burrow, the inclination of which is typically within 10° of the bedding planes of the sediment. These burrows can be very large, over a meter long in sediments that show good preservation, e.g. Jurassic rocks of the Yorkshire Coast (eastern United Kingdom), but the width is usually only up to 2 cm, restricted by the size of the organisms producing it. It is thought that they represent fodinichnia as the animal (probably a polychaete) scoured the sediment for food. Ichnogenus The ichnogenus ''Rhizocorallium'' Zenker 1836 includes three ichnospecies: ''Rhizocorallium jenense'' Zenker 1836 representing straight, short U-shaped spreite Spreite, meaning Lamina (leaf), leaf-blade in German (or spreiten, the plural form in German language, German) is a stacked, curved, layered structure that is characteristic of certain trace fossils. They are formed by invertebrate organisms tunn ...-burrows commonly oblique to bedding plane, and only rare ...
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Bulletin Of Geosciences
The ''Bulletin of Geosciences'' is a peer-reviewed scientific journal publishing original research papers, review articles, and short contributions. It covers all aspects of palaeoenvironmental geology, including palaeontology, stratigraphy, sedimentology, palaeogeography, palaeoecology, palaeoclimatology, geochemistry, mineralogy, geophysics, and related fields. It is published by the Czech Geological Survey, West Bohemian Museum in Plzeň, Palacký University Olomouc and the Geological institute of Czech Academy of Sciences. Abstracting and indexing This journal is abstracted and indexed in: * Science Citation Index Expanded * Current Contents/Physical, Chemical & Earth Sciences * Scopus * GeoRef The journal is included in the Geoscience e-Journals collection. According to the ''Journal Citation Reports'', the journal has a 2013 impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate tha ...
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Stratum
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as either '' bedding surfaces'' or ''bedding planes''.Salvador, A. ed., 1994. ''International stratigraphic guide: a guide to stratigraphic classification, terminology, and procedure. 2nd ed.'' Boulder, Colorado, The Geological Society of America, Inc., 215 pp. . Prior to the publication of the International Stratigraphic Guide, older publications have defined a stratum as either being either equivalent to a single bed or composed of a number of beds; as a layer greater than 1 cm in thickness and constituting a part of a bed; or a general term that includes both ''bed'' and ''lamina''.Neuendorf, K.K.E., Mehl, Jr., J.P., and Jackson, J.A. , eds., 2005. ''Glossary of Geology'' 5th ed. Alexandria, Virginia, American Geological Institute. 779 pp. . ...
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Ethology
Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objective study of animal behaviour, usually referring to measured responses to stimuli or to trained behavioural responses in a laboratory context, without a particular emphasis on evolutionary adaptivity. Throughout history, different naturalists have studied aspects of animal behaviour. Ethology has its scientific roots in the work of Charles Darwin and of American and German ornithologists of the late 19th and early 20th century, including Charles O. Whitman, Oskar Heinroth, and Wallace Craig. The modern discipline of ethology is generally considered to have begun during the 1930s with the work of Dutch biologist Nikolaas Tinbergen and Austrian biologists Konrad Lorenz and Karl von Frisch, the three recipients of the 1973 Nobel Prize in Phys ...
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