Dinichthys
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Dinichthys
''Dinichthys'' (from el, δεινός , 'terrible' and el, ἰχθύς 'fish') is an extinct monospecific genus of giant, marine arthrodire placoderm from the Late Devonian (Famennian stage), comparable in size, shape, and ecological role to the better-known ''Dunkleosteus''. Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio. Classification History ''Dinichthys'' was originally described in 1868 by John Newberry on the basis of an incomplete skull roof and mandibles (holotype AMNH 81). Subsequently, many unrelated large arthrodires were originally classified together within this genus, including species now assigned to ''Dunkleosteus'', '' Eastmanosteus'', and ''Titanichthys''. Notably, the type species of ''Dunkleosteus'' was originally described as ''Dinichthys terrelli'' by Newberry in 1873, and was later separated into ''Dunkleosteus'' by Jean-Pierre Lehman in 1956. ''Dunkleosteus'' was still thought to be close ...
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Dunkleosteus
''Dunkleosteus'' is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', and ''D. raveri''. The largest and most well known species is ''D. terrelli'', which grew up to long and in weight. ''Dunkleosteus'' could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of at the tip and at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco. Etymology ''Dunkleosteus'' was named in 1956 to honour David Dunkle (1911–1982), former curator of vertebrate paleontology at the Cleveland Museum of Natural History. The genus name ''Dunkleosteus'' combines David Dunkle's sur ...
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Aspinothoracidi
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator ''Dinichthys'', is the best-known member of this group. Many other genera, such as the infamous ''Dunkleosteus'', were previously thought to be close relatives of ''Dinichthys'' and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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Dunkleosteidae
Dunkleosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny While members of Dunkleosteidae were previously thought to be close relatives of the genus ''Dinichthys'' (when they were not synonymized as each other) and grouped together in the family Dinichthyidae, more recent phylogenetic studies have shown that the two taxa represent two very distinct clades within Arthrodira. Dunkleosteidae was then established as the sister taxon to the family Panxiosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). Dunkleosteidae was thus cladistically defined as including the type genus ''Dunkleosteus'' and all other genera in Dunkleosteoidea more closely related to ''Dunkleosteus'' than to '' Panxiosteus''. The phylogeny of Dunkleosteidae from the 2013 Zhu & Zhu study is shown in the ...
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Eastmanosteus
''Eastmanosteus'' ("Eastman's bone") is a fossil genus of dunkleosteid placoderms. It was closely related to the giant ''Dunkleosteus'', but differed from that genus in size, in possessing a distinctive tuberculated bone ornament, a differently shaped nuchal plate and a more zig-zagging course of the sutures of the skull roof.http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200812/W020090813371138329343.pdf Species of ''Eastmanosteus'' had powerful jaws with sharp cutting edges and were likely active predators. Fossils have been found in many parts of the world in marine sediments dating from the Middle to Late Devonian. They were medium-to-large fish, with specimens ''E. pustulosus'' and ''E. licharevi'' approaching a total length of 3 metres. Complete exoskeletons with soft-tissue traces of ''E. calliaspis'' from Australia make this one of the best known dunkleosteids. Phylogeny ''Eastmanosteus'' and its relative ''Dunkleosteus'' belong to the family Dunkleosteidae. The phylogeny of ...
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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus ...
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John Strong Newberry
John Strong Newberry (December 22, 1822 – December 7, 1892) was an American physician, geologist and paleontologist. He participated as a naturalist and surgeon on three expeditions to explore and survey the western United States. During the Civil War he served in the US Sanitary Commission and was appointed secretary of the western department of the commission. After the war he became professor of geology and paleontology at Columbia University School of Mines. Biography John Strong Newberry was born in Windsor, Connecticut to Henry and Elizabeth Strong. At the age of two he moved with his family to northeastern Ohio where his father opened a coal mining business. The fossils found in the coal deposits stimulated his interest in science and a visit in 1841 with James Hall, an eminent geologist and paleontologist, furthered his interests. He graduated from Western Reserve College in 1846 and from Cleveland Medical School in 1848. That same year he married Sarah Gaylord and ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
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Pachyosteomorphi
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References Arthrodires {{Placoderm-stub ...
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Jean-Pierre Lehman
Jean-Pierre Lehman (10 August 1914 – 26 February 1981) was a French paleontologist who specialized on tetrapods, particularly the actinopterygians. He followed early ideas comparative anatomy to study evolution through cladistic ideas and making use of biogeographical affinities. Lehman was born in Caen to Madeleine Auvray and civil engineer Gaston Lehman. Educated at Lycée Carnot, and the Sorbonne he also went to Grenoble where he met and married Ingegärd Eneström, daughter of Swedish physician Erik Hjalmar Eneström. Visits to Sweden brought him into contact with Erik Stensiö at the Museum of Natural History of Stockholm where he worked on the vertebrates of the Upper Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, wher ... of Scania in 1936–37. His thesis on the scales o ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Late Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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