Cyatheaceae
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Cyatheaceae
The Cyatheaceae are a family of ferns, the scaly tree ferns, one of eight families in the order Cyatheales in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may defined much more broadly (Cyatheaceae ''sensu lato'') as the only family in the Cyatheales, with the PPG I family treated as the subfamily Cyatheoideae. The narrower circumscription is used in this article. The family includes the world's tallest tree ferns, which reach heights up to 20 m. They are also very ancient plants, appearing in the fossil record in the late Jurassic, though the modern genera likely appeared in the Cenozoic. Cyatheaceae are the largest family of tree ferns, including about 640 species. Cyatheaceae and Dicksoniaceae, together with Metaxyaceae and Cibotiaceae, do not form a strongly supported monophyletic group and could be paraphyletic, but several individual subgroups are well supported as being monophyletic. Cyatheaceae are leptosporangiate ferns, the ...
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Cyatheales
The order Cyatheales, which includes most tree ferns, is a taxonomic order of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms from a vertical rhizome, others have shorter or horizontal expanding rhizomes. Some species have scales on the stems and leaves, while others have hairs. However, most plants in the Cyatheales are tree ferns and have trunk-like stems up to tall. It is unclear how many times the tree form has evolved and been lost in the order.Judd, W.S., C.S. Campbell, E.A. Kellogg, P.F. Stevens, and M.J. Donoghue (Eds.) 2008. Plant Systematics: A Phylogenetic Approach, Third Edition. Sinauer Associates, Sunderland, Massachusetts, USA. Description While the Cyatheales have been shown to be monophyletic through molecular analysis, no prominent morphological characteristics are common to the entire group ...
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Alsophila (plant)
''Alsophila'' is a genus of tree ferns in the family Cyatheaceae. It has also been considered to be a section in the subgenus ''Cyathea'' of the genus ''Cyathea''. Description Species of ''Alsophila'' have a treelike growth habit, with an erect trunk that rarely branches, or sometimes a more shrublike habit, with a creeping stem. Their fronds are large, with a strawlike stalk (stipe), dark brown or black in colour. Brown or dark brown scales are present, with distinct margins. The blade (lamina) of the frond is divided one to three times (one- to three-pinnate). The sori (spore-producing structures) are rounded and borne on smaller veins on the lower surface of the frond. An indusium (a covering to the sori) may or may not be present; if present initially, it may be lost as the frond ages. ''Alsophila'' is now separated from the other genera in the family Cyatheaceae primarily on the basis of molecular phylogenetic studies. The scales on the stalks (petioles) provide a morphol ...
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Sphaeropteris
''Sphaeropteris'' is a genus of tree fern in the family Cyatheaceae. It has been treated as a subgenus within the genus ''Cyathea'', but is accepted in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Description Species of ''Sphaeropteris'' have a treelike habit, usually with a single tall stem, and large fronds to or more in length. The stalk (stipe) of the frond is strawlike or purple in colour, with pale to brown scales. The sori (spore-bearing structures) are rounded, with or without indusia (covers). ''Sphaeropteris'' is now separated from the other genera in the family Cyatheaceae primarily on the basis of molecular phylogenetic studies. However, the scales on the stalks (petioles) provide a morphological distinction. ''Sphaeropteris'' has scales without distinct margins, whereas the other genera have scales with distinct margins. Taxonomy The genus ''Sphaeropteris'' was erected by Johann Jakob Bernhardi in 1801. It is placed in the family Cyatheacea ...
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Cyathea
''Cyathea'' is a genus of tree ferns, the type genus of the fern order Cyatheales. The genus name ''Cyathea'' is derived from the Greek ''kyatheion'', meaning "little cup", and refers to the cup-shaped sori on the underside of the fronds. Description The species of ''Cyathea'' are mostly terrestrial ferns, usually with a single tall stem. Rarely, the trunk may be branched or creeping. Many species also develop a fibrous mass of roots at the base of the trunk. The genus has a pantropical distribution, with over 470 species. They grow in habitats ranging from tropical rain forests to temperate woodlands. Classification Conant ''et al.'' in 1996, concluded on molecular cpDNA and morphological evidence that a system of three clades – ''Alsophila'', ''Cyathea'' and ''Sphaeropteris'' was the most accurate reflection of evolutionary lineages within the Cyatheaceae, ''Alsophila'' being the most basal and ''Cyathea'' and ''Sphaeropteris'' derived sister groups. In the Pteridophyte ...
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Gymnosphaera
Gymnosphaera is a genus of tree ferns in family Cyatheaceae. Taxonomy The genus was originally described by Carl Ludwig Blume in 1828. It was frequently treated by later authors as a synonym of ''Cyathea'' or '' Alsophila''. In the Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ... classification of 2016 (PPG I), ''Gymnosphaera'' was left as a synonym of ''Alsophila'', as evidence to support its reliable separation was lacking. More recently, Shi-Yong Dong and collaborators proposed the revival and recircumscription of the genus based on additional phylogenetic evidence, an act subsequently accepted by other tree fern workers. List of species , the following species are accepted in the genus as currently circumscribed by the Checklist of Ferns and Lycop ...
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Dicksoniaceae
Dicksoniaceae is a group of tropical, subtropical and warm temperate ferns, treated as a family in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), and counting 30-40 species. Alternatively, the family may be sunk into a very broadly defined family Cyatheaceae ''sensu lato'' as the subfamily Dicksonioideae. Most of the genera in the family are terrestrial ferns or have very short trunks compared to tree ferns of the family Cyatheaceae ''sensu stricto''. However, some of the larger species can reach several metres in height. A number of others are epiphytes. They are found mostly in tropical regions in the Southern Hemisphere, as far south as southern New Zealand. Larger tree ferns in the genus ''Cibotium'' were formerly included in Dicksoniaceae, but are now segregated as the family Cibotiaceae. Description Species in the family are generally characterized by large pinnate fronds, 1–4 m long. The family includes several species of tree ferns, which grow a single ...
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Cyathea Glauca
''Alsophila glaucifolia'', synonym ''Cyathea glauca'', is a species of tree fern endemic to Réunion. Little is known about this species. Habitat and related species ''A. glaucifolia'' grows at higher altitudes () and it is one of three species of tree fern that are indigenous to Réunion island. * '' Alsophila celsa'' (syn. ''Cyathea excelsa'') which also occurs in Mauritius, grows at slightly lower altitudes (200-1700m). Like ''A. glaucifolia'', its leaves are tripinnate, but the new fronds of ''A. celsa'' are scaleless; those of ''A. glaucifolia'' have red-brown scales. * '' Alsophila borbonica'' (syn. ''Cyathea borbonica'') is the only species with bipinnate fronds. Image:Cyathea glauca fronds.jpg, The fronds are tripinnate (branching to three levels). Image:Cyathea glauca.JPG, Growing under '' Acacia heterophylla'' in the forest of Bélouve, Réunion Image:Image-Cyathea glauca insertion of fronds.JPG, Detail of trunk and insertion of fronds. New growth is ...
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Polypodiophyta
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The Polypodiophyta, polypodiophytes include all living pteridophytes except the Lycopodiopsida, lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaf, leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled Fiddlehead fern, fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae (plant), Polypodiidae) and eusporangiate ferns, the latter group including horse ...
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Cibotiaceae
''Cibotium'' (from Greek , ''kibṓtion'', "little chest" or "box"), also known as manfern, is a genus of 11 species of tropical tree ferns. It is the only genus in family Cibotiaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may be treated as the subfamily Cibotioideae of a very broadly defined family Cyatheaceae, the family placement used for the genus in '' Plants of the World Online'' . Species , '' Plants of the World Online'' accepted the following species and hybrids: Some extinct species have also been placed in this genus: *†''Cibotium iwatense'' Ogura *†''Cibotium oregonense'' Barrington Distribution Species of the genus are distributed fairly narrowly in Hawaii (four species, plus a hybrid, collectively known as ''hāpuu''), Southeast Asia (five species), and the cloud forests of Central America and Mexico (two species). The natural habitat of ''Cibotium'' is among the dripping trees and stream gullies of the ra ...
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Georg Friedrich Kaulfuss
Georg Friedrich Kaulfuss (8 April 1786 – 9 December 1830) was a professor at Halle. He described the pteridophytes collected by Adelbert von Chamisso, and he named the fern ''Cibotium chamissoi'' after him. The genus ''Kaulfussia'' is named for Kaulfuss. In 1816 he obtained his doctorate from the University of Halle, where in 1823 he became a full professor of botany. Plants Named by Kaulfuss More than 200 plant names were published by Kaulfuss.Kaulfuss, Georg Friedrich (names)
Tropicos.org. Missouri Botanical Garden. 01 Feb 2012 These include the genera: * *'''' * *'' Balantium'' * Plant families named by Kaulfuss in ...
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Pteridophyte Phylogeny Group
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relationships discovered through phylogenetic studies. In 2016, the group published a classification for extant pteridophytes, termed "PPG I". The paper had 94 authors (26 principal and 68 additional). PPG I A first classification, PPG I, was produced in 2016, covering only extant (living) pteridophytes. The classification was rank-based, using the ranks of class, subclass, order, suborder, family, subfamily and genus. Phylogeny The classification was based on a consensus phylogeny, shown below to the level of order. The very large order Polypodiales was divided into two suborders, as well as families not placed in a suborder: Classification to subfamily level To the level of subfamily, the PPG I classification is as follows. *Class Lycopodi ...
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Metaxyaceae
''Metaxya'' is a neotropical genus of ferns in the order Cyatheales. It is the only genus in the family Metaxyaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed in the subfamily Metaxyoideae of a more broadly defined family Cyatheaceae, the family placement used in ''Plants of the World Online'' . The species of the genus are characterized by large fronds that approach 8 ft (2.5 m) in length. Species , ''Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...'' accepted the following species: References External links * * Cyatheales Fern genera {{Cyatheales-stub ...
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