Culbersonia
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Culbersonia
''Culbersonia'' is a fungal genus in the family Caliciaceae. This is a monotypic genus, containing the single foliose lichen ''Culbersonia nubila'' (formerly called ''Culbersonia americana''). This species, which grows on trees and rocks, is found in dry subtropical regions of the world, particularly in Africa and Central America. Taxonomy The genus ''Culbersonia'' was circumscribed by Theodore Esslinger in 2000, to contain a lichen he collected from Arizona that he named ''Culbersonia americana''. The genus name of ''Culbersonia'' is in honour of Chicita Frances 'Nan' Culberson, (b.1931), an American botanist (Mycology und Lichenology), who worked as a botanical researcher at Duke University in Durham/North Carolina and William Louis "Bill" Culberson (1929–2003), an American lichenologist. As they were "longtime friends and mentors" of Esslinger. After ''Culbersonia americana'' was published, it was determined that the Arizona species was the same as one previously describ ...
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Caliciaceae
The Caliciaceae are a family of mostly lichen-forming fungi belonging to the class Lecanoromycetes in the division Ascomycota. Although the family has had its classification changed several times throughout its taxonomic history, the use of modern molecular phylogenetic methods have helped to establish its current placement in the order Caliciales. Caliciaceae contains 36 genera and about 600 species. The largest genus is ''Buellia'', with around 300 species; there are more than a dozen genera that contain only a single species. Most Caliciaceae grow on bark, dead wood, or rocks. Some members of this family, particularly those of the type genus, ''Calicium'', are characterized by the presence of thin-walled and short-lasting asci (spore-bearing cells) and a mazaedium, which is an accumulation of loose, maturing spores covering the surface of the fruiting body. The resulting passive spore dispersal is relatively rare amongst the Ascomycota. The mazaedium, usually black, either ...
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Physciaceae
The Physciaceae are a family of mostly lichen-forming fungi belonging to the class Lecanoromycetes in the division Ascomycota. A 2016 estimate placed 19 genera and 601 species in the family. Genera This is a list of the genera contained within the Physciaceae, based on a 2020 review and summary of ascomycete classification. Following the genus name is the taxonomic authority, year of publication, and the number of species: *''Anaptychia'' – about 15 spp. *'' Coscinocladium'' – 2 spp. *''Heterodermia'' – about 90 spp. *''Hyperphyscia'' – 9 spp. *'' Kashiwadia'' – 1 sp. *''Leucodermia'' – 10 spp. *'' Mischoblastia'' – 3 spp. *'' Mobergia'' – 1 sp. *''Oxnerella'' – 1 spp. *'' Phaeophyscia'' – 66 spp. *''Phaeorrhiza'' – 2 spp. *''Physcia'' – about 80 spp. *'' Physciella'' – 4 spp. *''Physconia'' – about 25 spp. *'' Polyblastidium'' – 18 spp. *''Rinodina'' – about 300 spp *'' Rinodinella'' – 6 spp. *'' Tornabea'' – 1 spp. The ge ...
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William Louis Culberson
William Louis "Bill" Culberson (April 5, 1929 in Indianapolis, Indiana – February 8, 2003 in Durham, North Carolina) was an American lichenologist. Professional history Culberson earned his bachelor's degree at the University of Cincinnati, where he was influenced by E. Lucy Braun; he subsequently attended the University of Paris and the University of Wisconsin–Madison. In 1955, Culberson joined the botany department at Duke University; he subsequently managed Duke's acquisition of the lichen-centric herbaria of Julien Harmand and Johan Havaas. He served as the Hugo L. Blomquist Professor. In 2010, the lichen collection was officially named the William Louis & Chicita F. Culberson Lichen Herbarium & Library. He served as president of the Botanical Society of America and the American Bryological and Lichenological Society and as director of the Sarah P. Duke Gardens. He was the first editor-in-chief of the journal ''Systematic Botany''. In 1992, he became one of the f ...
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Caliciales Genera
Caliciales is an order of mostly lichenized fungi in the class Lecanoromycetes. It consists of two families: Caliciaceae and Physciaceae, which together contain 54 genera and more than 1200 species. The order was circumscribed by American botanist Charles Edwin Bessey in 1907. Families and genera , Species Fungorum (in the Catalogue of Life) accepts 2 families, 56 genera, and 910 species in the Caliciales. *Caliciaceae ::''Acolium'' – 5 spp. ::'' Acroscyphus'' – 1 sp. ::''Allocalicium'' – 1 sp. ::''Amandinea'' – 83 spp. ::''Australiaena'' – 1 sp. ::''Baculifera'' – 18 spp. ::''Buellia'' – 201 spp. ::'' Caliciella'' – 1 sp. ::''Calicium'' – 36 spp. ::'' Chrismofulvea'' – 3 spp. ::'' Ciposia'' – 1 sp. ::'' Cratiria'' – 23 spp. ::'' Dermatiscum'' – 2 sp. ::'' Dermiscellum'' – 1 sp. ::'' Dimelaena'' – 10 spp. ::'' Diploicia'' – 6 spp. ::'' Diplotomma'' – 12 spp. ::'' Dirinaria'' – 18 spp. ::'' Endohyalina'' – 10 sp. ::'' Fluctua'' –  ...
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Caliciales
Caliciales is an order of mostly lichenized fungi in the class Lecanoromycetes. It consists of two families: Caliciaceae and Physciaceae, which together contain 54 genera and more than 1200 species. The order was circumscribed by American botanist Charles Edwin Bessey in 1907. Families and genera , Species Fungorum (in the Catalogue of Life) accepts 2 families, 56 genera, and 910 species in the Caliciales. *Caliciaceae ::''Acolium'' – 5 spp. ::'' Acroscyphus'' – 1 sp. ::''Allocalicium'' – 1 sp. ::''Amandinea'' – 83 spp. ::''Australiaena'' – 1 sp. ::''Baculifera'' – 18 spp. ::''Buellia'' – 201 spp. ::'' Caliciella'' – 1 sp. ::''Calicium'' – 36 spp. ::''Chrismofulvea'' – 3 spp. ::''Ciposia'' – 1 sp. ::''Cratiria'' – 23 spp. ::''Dermatiscum'' – 2 sp. ::''Dermiscellum'' – 1 sp. ::''Dimelaena'' – 10 spp. ::''Diploicia'' – 6 spp. ::''Diplotomma'' – 12 spp. ::''Dirinaria'' – 18 spp. ::''Endohyalina'' – 10 sp. ::'' Fluctua'' – 1 sp. ::''Gas ...
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Lichen Genera
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Ascospore
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Diplotomma
''Diplotomma'' is a genus of lichenized fungi in the family Caliciaceae. The genus has a widespread distribution and contains about 29 species. The genus was circumscribed by Julius von Flotow in 1849. It was later wrapped into ''Buellia'' before being segregated from that genus by David Hawksworth in 1980. Selected species , Species Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and the Institute of M ... accepts 10 species of ''Diplotomma'': *'' Diplotomma alboatrum'' *'' Diplotomma cedricola'' *'' Diplotomma epipolium'' *'' Diplotomma glaucoatrum'' *'' Diplotomma hedinii'' *'' Diplotomma murorum'' *'' Diplotomma parasiticum'' *'' Diplotomma pharcidium'' *'' Diplotomma venustum'' *'' Diplotomma vezdanum'' References Caliciales Lichen genera Caliciales genera Taxa described ...
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Pyxine
''Pyxine'' is a genus of foliose lichens in the family Caliciaceae. The genus has a widespread distribution in tropical regions. The presence or absence of the compound lichexanthone is a character Character or Characters may refer to: Arts, entertainment, and media Literature * ''Character'' (novel), a 1936 Dutch novel by Ferdinand Bordewijk * ''Characters'' (Theophrastus), a classical Greek set of character sketches attributed to The ... used in classifying ''Pyxine '' species; about 20 species contain this compound. This represents the largest group of foliose lichens with the compound. Species *'' Pyxine albovirens'' *'' Pyxine astipitata'' – Brazil *'' Pyxine astridiana'' – neotropical *'' Pyxine australiensis'' – Asia; Australia *'' Pyxine berteriana'' – pantropical *'' Pyxine boonpragobiana'' – Thailand *'' Pyxine coccifera'' *'' Pyxine cocoes'' *'' Pyxine cognata'' - pantropical *'' Pyxine consocians'' *'' Pyxine convexior'' *'' Pyxine copela ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Molecular Phylogenetic
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical framew ...
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