Ctenochasmatoidea
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Ctenochasmatoidea
Ctenochasmatoidea is a group of early pterosaurs within the suborder Pterodactyloidea. Their remains are usually found in what were once coastal or lake environments. They generally had long wings, long necks, and highly specialized teeth. Evolutionary history The earliest known ctenochasmatoid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Ecology Most ctenochasmatoids were aquatic or semi-aquatic pterosaurs, possessing large webbed hindfeet and long ...
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Euctenochasmatia
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of ''Pterodactylus'' and ''Ctenochasma'', and all their descendants. Euctenochasmatians were specialized pterosaurs that had elongated necks as well as specialized teeth. A peculiar family within this group is the Ctenochasmatidae, which most of the members had very distinguishing teeth that were lined within their elongated snouts. A genus called ''Pterofiltrus'' only had 112 teeth, but these teeth cover about 55.8% of the total skull, and the skull itself measured about in length. Description Euctenochasmatians had very distinctive features in comparison to other pterosaurs, including the shape of their jaws, as well as their highly specialized teeth. These teeth are thought to have been used for filter-feeding, the genus ''Pterodaustro'' for example, had a long snout and its lower jaws curve strongly upwards, and th ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like ''Nyctosaurus'' and ''Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. "Pteroda ...
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Gallodactylidae
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey. History Gallodactylidae was named to contain ''Gallodactylus'' (now usually considered a synonym of ''Cycnorhamphus'') and its closest relatives. Many subsequent studies, however, showed that ''Gallodactylus'' did not form a clade with any non-synonymous pterosaurs that were not themselves part of a different family, and so the name was often ignored. The name returned to common use with the discovery of ''Gladocephaloideus'', a Chinese pterosaur species that shared many similarities with ''Cycnorhamphus''. Among other features, the Gallodactylidae was distinguished by ...
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Pterodactylus
''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of ''Pterodactylus'' have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of ''Pterodactylus'' have tentatively been identified from elsewhere in Europe and in Africa. ''Pterodactylus'' was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, ''Pterodactylus'' had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. ''Pterodactylus'' was a small pterosaur compared to other famo ...
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Gallodactylid
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey. History Gallodactylidae was named to contain ''Gallodactylus'' (now usually considered a synonym of ''Cycnorhamphus'') and its closest relatives. Many subsequent studies, however, showed that ''Gallodactylus'' did not form a clade with any non-synonymous pterosaurs that were not themselves part of a different family, and so the name was often ignored. The name returned to common use with the discovery of ''Gladocephaloideus'', a Chinese pterosaur species that shared many similarities with ''Cycnorhamphus''. Among other features, the Gallodactylidae was distinguished by ...
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Diopecephalus
''Diopecephalus'' is a genus of pterodactyloid pterosaur from the Lower Tithonian (Upper Jurassic) of the Lithographic Limestone, Bavaria, Germany. The type and only species is ''D. kochi'', although the name has been applied to ''Pterodactylus longicollum'', with ''longicollum'' erroneously listed as the type species. Assigned species Like many pterosaurs, it has had a confusing taxonomic history, being given names by various authorities which identify it with four other genera: *''Pterodactylus longicollum'' (von Meyer 1854) *''Pterodactylus vulturinus'' (Wagner 1857) *''Pterodactylus longicollis'' (Wagner 1858) *''Pterodactylus suevicus'' (Fraas 1878) *''Cycnorhamphus fraasii'' (Seeley 1891) *''Pterodactylus frassi'' (Seeley 1901) *''Gallodactylus longicollum'' (Fabre 1974) *''Ornithocephalus longipes'' (Olshevsky 1978) (''Ornithocephalus'' is an obsolete name for ''Pterodactylus'') *''Ornithocephalus vulturinus'' (Olshevsky 1978) In 2017, Steven U. Vidovic and David M. Mart ...
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Gladocephaloideus Jingangshanensis
''Gladocephaloideus'' is a genus of ctenochasmatid ctenochasmatoid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now western Liaoning, China. Discovery ''Gladocephaloideus'' is known from a complete skull and partial postcranial skeleton including traces of hair-like pycnofibres. It was collected from the Jingangshan Bed of the Yixian Formation, dating to the early Aptian stage, about 121 million years ago. It was named by Lü Junchang, Ji Qiang, Wei Xuefang and Liu Yongqing in 2012 and the type species is ''Gladocephaloideus jingangshanensis''. The generic name is derived from Latin ''gladius'', "sword", and Greek κεφαλή, ''kephalè'', "head", en εἶδος, ''eidos'', "form", referring to the elongated form of the skull. The specific name refers to the provenance from the Jiangangshan. The holotype, IG-CAGS-08-07, includes the almost complete skull, the lower jaws, the front neck vertebrae and part of a hindlimb. In 2016, a second speci ...
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Cycnorhamphus
''Cycnorhamphus'' (meaning "swan beak") is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. It is probably synonymous with the genus ''Gallodactylus''. History In 1855, a fossil in a plate of shale from the Kimmeridgian, found near Nusplingen in Württemberg, holotype GPIT "Orig. Quenstedt 1855, Taf. 1" or GPIT 80, was named ''Pterodactylus Suevicus'' by Friedrich August Quenstedt. The specific name refers to the tribal area of Suevia. Quenstedt had earlier mentioned the find in a letter to Professor Heinrich Georg Bronn, which was published in 1854. In it he used the name ''Pterodactylus Württembergicus''. In 1855 and 1858, Christian Erich Hermann von Meyer adopted this older species name but it would be forgotten afterwards. The publication in 1854 was not meant to be a nomenclatural act. According to Peter Wellnhofer, ''Pterodactylus württembergicus'' should be considered a ''nomen oblitum' ...
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Filter-feeding
Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feeding are clams, krill, sponges, baleen whales, and many fish (including some sharks). Some birds, such as flamingos and certain species of duck, are also filter feeders. Filter feeders can play an important role in clarifying water, and are therefore considered ecosystem engineers. They are also important in bioaccumulation and, as a result, as indicator organisms. Fish Most forage fish are filter feeders. For example, the Atlantic menhaden, a type of herring, lives on plankton caught in midwater. Adult menhaden can filter up to four gallons of water a minute and play an important role in clarifying ocean water. They are also a natural check to the deadly red tide. Extensive article on the role of menhaden in the ecosystem and possible resul ...
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Ardeadactylus
''Ardeadactylus'' (from ''Ardea'' – meaning "heron", and also a name of a genus of herons – and ''dactylus'', meaning "finger") is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic Solnhofen limestone of Bavaria, southern Germany. It contains a single species, ''Ardeadactylus longicollum'', which was originally thought to be a species of ''Pterodactylus'', as ''P. longicollum''. History Only two specimens of ''Ardeadactylus'' are known to exist currently: SMNS 56603 (earlier SMNS 5802) found in 1874, a specimen from Nusplingen initially thought to belong to the species '' Pterodactylus suevicus'' (currently ''Cycnorhamphus''), and the neotype of the species, JME-SOS 2428, a specimen held at Jura Museum in Eichstätt. Other known specimens, including the holotype designated by Christian Erich Hermann von Meyer when he named the type species ''Pterodactylus longicollum'' in 1854,Meyer, C.E.H. von, 1854, "Mittheilungen an Professor Bronn: Anthracotheri ...
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Aurorazhdarcho
''Aurorazhdarcho'' is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic period (early Tithonian stage) of what is now Bavaria, southern Germany. History First discovery A specimen, originally classified as ''Pterodactylus micronyx'' (now ''Aurorazhdarcho micronyx''), was one of the earliest, if not the earliest, documented pterosaur fossil ever found. The holotype specimen of ''P. micronyx'', also known as the "Pester Exemplar", was originally part of the private fossil collection held by Archduchess Maria Anna of Austria. Evidence suggest that the Pester Exemplar was unearthed at some point between 1757, when Maria Anna was recovering from serious cases of pneumonia and tuberculosis, after which she began collecting fossils, and 1779, when the specimen was first studied by scientists. This overlaps with the possible time of discovery of the holotype specimen of ''Pterodactylus antiquus'', often considered the first pterosaur found, which was unearthe ...
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