Craniformea
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Craniformea
Craniata is a class of brachiopods originating in the Cambrian period and still extant today. It is the only class within the subphylum Craniiformea, one of three major subphyla of brachiopods alongside linguliforms and rhynchonelliforms. Craniata is divided into three orders: the extinct Craniopsida and Trimerellida, and the living Craniida, which provides most information on their biology. Living members of the class have shells which are composed of calcite, though some extinct forms my have aragonite shells. The shells are inarticulate (lack a hinge with distinct tooth-and-socket connections) and are usually rounded in outline. There is no pedicle, with the rear edge of the body cavity having the form of a smooth and flat wall perforated by the anus. This class of brachiopods has an unsupported lophophore with only a single row of tentacles. In the absence of a pedicle, the shell is usually attached directly to a hard substrate. Many craniiforms are encrusting animals whic ...
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Rhynchonelliformea
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The name, Rhynchoelliformea, which replaces Articulata, which is also a class of crinoids, comes from the family Rhynchonellidae, which however is no more representative of articulate brachiopods than any other group such as spirifers or strophomenids. It just happens to be the name chosen, based on an included taxon. The main difference between the Rhynchonelliformea described in the Treatise Part H, revised 2000/2007, and the Articulata of the Treatise part H, 1965, lies in the groups included, their taxonomic po ...
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Subphylum
In zoological nomenclature, a subphylum is a taxonomic rank below the rank of phylum. The taxonomic rank of "subdivision (rank), subdivision" in fungi and plant taxonomy is equivalent to "subphylum" in zoological taxonomy. Some plant taxonomists have also used the rank of subphylum, for instance monocotyledons as a subphylum of phylum Angiospermae and vertebrates as a subphylum of phylum Chordata. Taxonomic rank Subphylum is: #subordinate to the phylum #superordinate to the infraphylum. Where convenient, subphyla in turn can be divided into infraphyla; in turn such an infraphylum also would be superordinate to any classes or superclasses in the hierarchy. Examples Not all fauna phyla are divided into subphyla. Those that are include: *Arthropoda: divided into subphyla Trilobitomorpha, Chelicerata, Myriapoda, Hexapoda and Crustacean, Crustacea, *Brachiopoda: divided into subphyla Linguliformea, Craniformea and Rhynchonelliformea, *Chordata: divided into Tunicate, Tunicata, ...
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Linguliformea
Linguliformea is a subphylum of inarticulate brachiopods. These were the earliest of brachiopods, ranging from the Cambrian into the Holocene. They rapidly diversified during the Cambrian into the Ordovician, but most families became extinct by the end of the Devonian. The articulation in these brachiopods is lacking. These brachiopods have adductor and oblique muscles, but no diductor muscles. The anus is located at the side of the body. The pedicle is a hollow extension of the ventral body wall. Posterior body wall separates dorsal and ventral mantles. The shells are usually made up of apatite (calcium phosphate), however rare cases have calcite or aragonite Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including prec ... shells. See also * References Brachiopod taxonomy Paleozoic ...
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Adductor Muscles (bivalve)
The adductor muscles are the main muscular system in bivalve mollusks (e.g. in clams, scallops, mussels, oysters, etc.). In many parts of the world, when people eat scallops, the adductor muscles are the only part of the animal which is eaten. Adductor muscles leave noticeable scars or marks on the interior of the shell's valves. Those marks (known as adductor muscle scars) are often used by scientists who are in the process of identifying empty shells to determine their correct taxonomic placement. Bivalve mollusks generally have either one or two adductor muscles. The muscles are strong enough to close the valves of the shell when they contract, and they are what enable the animal to close its valves tightly when necessary, such as when the bivalve is exposed to the air by low water levels, or when it is attacked by a predator. Most bivalve species have two adductor muscles, which are located on the anterior and posterior sides of the body. Some families of bivalves have on ...
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Lophophore
The lophophore () is a characteristic feeding organ possessed by four major groups of animals: the Brachiopoda, Bryozoa, Hyolitha, and Phoronida, which collectively constitute the protostome group Lophophorata.Introduction to the Lophotrochozoa
– Retrieved 3 May 2010
All lophophores are found in aquatic organisms.


Etymology

''Lophophore'' is derived from the Greek ''lophos'' (crest, tuft) and ''-phore'', ''-phoros'' (φορος) (bearing), a derivative of ''phérein'' (φέρειν) (to bear); thus crest-bearing.


Characteristics

The lophophore can most easily be described as a ring of ted tentacles surrounding the mouth, but it is often horseshoe-shape ...
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Isocrania Costata Sowerby 1823
''Isocrania'' is an extinct genus of brachiopods found during the Upper Cretaceous. Early representatives were attached to the underground, but later species are presumed to be free living at an increasingly earlier age. This was probably an adaptation to the increasing very thick and fine sedimentation during the latest Cretaceous. File:Crania ignabergensis lateral.jpg, ''Isocrania egnabergensis'', lateral view File:Crania ignabergensis internal.jpg, internal Description ''Isocrania'' is round to ovate, up to 1 cm in diameter, and has 15-65 strong ribs, that start at ±½mm from the origin of growth (or umbo). These ribs may extend beyond the edge of the valves. The umbo is not precisely in the centre of the valve. The attachment area is smaller than usual, and virtually absent in adults of later species. The dorsal valve is conical, the ventral valve flat to conical, flatter for adolescents and earlier species. The inner edge of the valves is flattened and grainy. Sp ...
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Anus
The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, depending on the type of animal, includes: matter which the animal cannot digest, such as bones; Summary at food material after the nutrients have been extracted, for example cellulose or lignin; ingested matter which would be toxic if it remained in the digestive tract; and dead or excess gut bacteria and other endosymbionts. Amphibians, reptiles, and birds use the same orifice (known as the cloaca) for excreting liquid and solid wastes, for copulation and egg-laying. Monotreme mammals also have a cloaca, which is thought to be a feature inherited from the earliest amniotes via the therapsids. Marsupials have a single orifice for excreting both solids and liquids and, in females, a separate vagina for reproduction. Female placental mamm ...
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Inarticulata
Inarticulata was historically defined as one of the two classes of the phylum Brachiopoda and referred to those having no hinge. The other class was Articulata, meaning articulated — having a hinge between the dorsal and ventral valves. These classifications have now been superseded, see brachiopod classification. See also * Treatise on Invertebrate Paleontology The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and co ... — also mentions the new Brachiopoda classification system References {{reflist Animal classes Obsolete animal taxa Brachiopod taxonomy ...
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Aragonite
Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including precipitation from marine and freshwater environments. The crystal lattice of aragonite differs from that of calcite, resulting in a different crystal shape, an orthorhombic crystal system with acicular crystal. Repeated twinning results in pseudo-hexagonal forms. Aragonite may be columnar or fibrous, occasionally in branching helictitic forms called ''flos-ferri'' ("flowers of iron") from their association with the ores at the Carinthian iron mines. Occurrence The type location for aragonite is Molina de Aragón in the Province of Guadalajara in Castilla-La Mancha, Spain, for which it was named in 1797. Aragonite is found in this locality as cyclic twins inside gypsum and marls of the Keuper facies of the Triassic. This type of arago ...
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Calcite
Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on the Mohs scale of mineral hardness, based on Scratch hardness, scratch hardness comparison. Large calcite crystals are used in optical equipment, and limestone composed mostly of calcite has numerous uses. Other polymorphs of calcium carbonate are the minerals aragonite and vaterite. Aragonite will change to calcite over timescales of days or less at temperatures exceeding 300 °C, and vaterite is even less stable. Etymology Calcite is derived from the German ''Calcit'', a term from the 19th century that came from the Latin word for Lime (material), lime, ''calx'' (genitive calcis) with the suffix "-ite" used to name minerals. It is thus etymologically related to chalk. When applied by archaeology, archaeologists and stone trade pr ...
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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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